New and noteworthy records of Plants, Lichens and Insects in Altai Territory and Republic of Altai (Southern Siberia). II.

Corresponding author: Roman Yakovlev (yakovlev_asu@mail.ru)

Academic editor: A. Matsyura | Received: 1 December 2025 | Accepted: 19 December 2025 | Published: 21 December 2025

Citation: Davydov EA, Kosachev P, Golyakov P, Zalutsky T, Svirin E, Kudrov O, Pavlova P, Slepchenko A, Knyazev S, Storozhenko Yu, Yakovchenko L, Yakovlev R (2025) New and noteworthy records of Plants, Lichens and Insects in Altai Territory and Republic of Altai (Southern Siberia). II. Acta Biologica Sibirica 11: 1499–1531. https://doi.org/10.5281/zenodo.18007146

Zoobank: http://zoobank.org/DD9A6A15-577D-4E87-81B5-9258FAE8B306

The present material continues the series of publications on the biodiversity of the Altai Territory and the Republic of Altai (Davydov et al. 2022, 2023; Kosachev and Albach 2022) and represents new records on the distribution of a number of species of vascular plants, lichens, and insects in the Altai Territory and the Republic of Altai, including species requiring special protection. While not all species documented in this study are currently designated for special protection, the identification of new or historically underrepresented populations of rare taxa underscores the critical need for up-to-date distributional data. Accurate and comprehensive information on the geographical spread and ecological preferences of species, particularly those with limited distributions or recognized conservation concern, is fundamental for their effective protection. This knowledge is essential for identifying priority areas for conservation, developing targeted protection strategies, and making informed decisions regarding their inclusion or status within regional and national Red Books. The present work aims to contribute to this vital effort by documenting new occurrences and ecological parameters for some vulnerable species, thereby providing essential evidence for conservation planning and enabling conservation efforts to be more precise and impactful.

The paper contains data obtained from the materials collected using traditional methods in various localities of the Altai Territory and the Republic of Altai.

The material is kept in the following collections:

ALTB – the herbarium of Altai State University (Barnaul) and its unit TIGZ – the herbarium of Tigirek Natural Reserve (Barnaul);

ESB – private collection of Egor Svirin (Barnaul); OKB – private collection of Oleg Kudrov (Barnaul);

RYB – research collection of Roman Yakovlev (Barnaul); TZB – private collection of Timofey Zalutsky (Barnaul).

Plants

During 2023, significant floristic discoveries were made within the Altai Territory. In a challenging-to-access sector of the Tigirek Nature Reserve (Beloretsk Floristic District), situated in the Belaya River valley, two species belonging to the genus Veronica (V. × sessiliflora and V. × schmakovii) were documented for the first time for the region's flora. Concurrently, the northern part of Altai Territory yielded two additional new species for the regional flora from the family Cyperaceae (Rhynchospora alba and Schoenoplectiella supina). Moreover, analysis of collected specimens and corresponding data from iNaturalist.org revealed the presence of four plant species previously considered rare within the Altai Territory: Gymnocarpium continentale, Scheuchzeria palustris, Cyperus michelianus, and Drosera rotundifolia. Three more new records were discovered during a study of coastal aquatic habitats in the plain part of the Altai Territory.

Family Cystopteridaceae

Gymnocarpium continentale (Petrov) Pojark.

Material examined. Russia, Altai Territory, Charyshskii District, upper reaches of the Kholodny Klyuch stream, a tributary of the Bolshoi Tigirek River, 51°04ʹ47.48ʺ N, 83°00ʹ05.23ʺ E, elev. 1264 m, cracks in granite rocks, 07.vii.2021, P. Kosachev (www.inaturalist.org/observations/96068979) (Fig. 1a, b); in the same place. 16.vii.2020. P.V. Golyakov (www.inaturalist.org/observations/53705977).

Distribution. Circumboreal species.

Note. The species is rare in the Altai Mountain Country; in the Altai Territory only one location is known on the Sinyukha Mt. (Kurinsky District) (Shmakov 2005).

Family Plantaginaceae

Veronica × schmakovii Kosachev

Material examined. Russia, Altai Territory, Zmeinogorsky District, the basin of the Belaya River, the valley of the Strizhanka River, 50°56'17.54ʺ N, 82°54'53.03ʺ E, elev. 630 m, bushy slope with southern exposure, rocks, 16.vii.2023, leg. P. Kosachev (TIGZ; www.inaturalist.org/observations/175151992; Fig. 2a–d) Russia, Altai Territory, Zmeinogorsky District, valley of the Belaya River near the mouth of the Krakhalikha River, 50°57ʹ25.3ʺ N, 82°51ʹ51.0ʺ E, elev. 576 m, floodplain meadow, bushy mountain slope, 19.vii.2023, leg. P. Kosachev (TIGZ; www.inaturalist.org/observations/175487299; Fig. 3a–c).

Distribution. This hybrid is endemic to the Altai Mountain Country.

Note. This is the first record of the species in the Altai Territory. V. × schmakovii previously known from 23 locations from the Republic of Altai, Republic of Tuva, Kazakhstan and Mongolia (Kosachev and Albach 2022).

Veronica × sessiliflora Bunge

Material examined. Russia, Altai Territory, Zmeinogorsky District, valley of the Belaya River near the mouth of the Krakhalikha River, 50°57ʹ25.3ʺ N, 82°51ʹ51.0ʺ E, elev. 576 m, floodplain meadow, bushy mountain slope, 19.vii.2023, leg. P. Kosachev (TIGZ; www.inaturalist.org/observations/175491652; Fig. 4a, b).

Distribution. Endemic to the Altai Mountain Country.

Note. This is the first record of the species in the Altai Territory. In total, 21 locations were known within the Altai Mts. (Kosachev and Albach 2022).

Family Scheuchzeriaceae

Scheuchzeria palustris L.

Material examined. Russia, Altai Territory, Talmensky District, 53°50ʹ11.68ʺ N, 83°46ʹ47.06ʺ E. Sphagnum bog with Andromeda polifolia L. and Oxycoccus palustris Pers., 29.x.2023, leg. P.V. Golyakov (www.inaturalist.org/observations/189323785); Altai Territory, Talmensky District, near the community of Vypolzovo, Klyukvennoye Lake, 53°50ʹ11.6ʺ N, 83°46ʹ46.1ʺ E, cranberry-sphagnum bog, 01.vii.2025, leg. P.V. Golyakov (ALTB, TIGZ).

Distribution. Holarctic plant. The species grows in sphagnum bogs in the forest zone.

Note. This is the second record of the species in the region. The first report was made by Silantyeva (2013) for the Pervomaysky District near the village of Bayunovskie Klyuchi on the Klyukvennoye bog. This is a rare plant in Russia, listed in some regional Red Books in the European part (Red Books of Tambov, Lipetsk, Penza, Voronezh, Moscow regions, the Republic of Mordovia and some others) (Red Book... 2019). The limiting factor for the distribution of the species in the southern regions is the limited number of biotopes – sphagnum bogs. Such habitats are typical for more northern regions of Siberia, and in the Altai Territory they are known only in the northern regions. In the Republic of Altai, the plant has also been collected from a site on the Lebed' River (Krasnoborov and Korotkova 1988; Gerasimovich 2012).

Due to the insignificant area of sphagnum bogs and their degradation, the only location of the plant in the region, we recommend a species for inclusion in the Red Book of the Altai Territory.

Family Cyperaceae

Cyperus michelianus (L.) Delile

Material examined. Russia, Altai Territory, Kalmanskii District, settlement Buranovo, bank of the Obi River, 53°2′33″ N, 83°37′35″ E, sandbank, 23.viii.2024, leg. P.V. Golyakov (ALTB; www.inaturalist.org/observations/237493344).

Distribution. Eurasia.

Note. A rare species in Siberia, where it is known only from 6 regions: Omsk, Tomsk, Novosibirsk, Chuta, Republic of Buryatia, and Altai Territory. In Novosibirsk Region, the species is listed in the Red Book (Artemov 2018). In Altai Territory, the plant was collected very rarely: in the floodplain of the Chumysh River (Talmensky District), Lake Shirokoe (Zarinsky District), the floodplain of the Obi River (Pervomaysky District), the vicinity of the village of Stan-Bekhtemir (Biysky District) (Silantieva 2013). The present record supplements the distribution of this rare species in Altai Territory.

Rhynchospora alba (L.) Vahl

Material examined. Russia, Altai Territory, Talmensky District, near the village of Vypolzovo, Klyukvennoye Lake, 53°50ʹ11.6ʺ N, 83°46ʹ46.1ʺ E, cranberry-sphagnum bog, 01.vii.2025, leg. P.V. Golyakov (ALTB).

Distribution. This species is widely distributed across the temperate zone of the Northern Hemisphere, with a preference for its southern regions, and is considered a circumpolar bog inhabitant.

Note. This represents the first record of the species in Altai Territory. In Siberia, it grows on Sphagnum or occasionally Hypnum bogs, at the edges of overgrowing lakes, and in waterlogged forests. It is found in eight Siberian regions: Tyumen, Omsk, Tomsk, and Irkutsk regions, Republic of Buryatia, Krasnoyarsk and Trans-Baikal Territories, and Yakutia (Timokhina and Bondareva 1990).

Schoenoplectiella supina (L.) Lye

Material examined. Russia, Altai Territory, near the city of Barnaul, Obi River floodplain, 53°15′53″ N, 83°52′13″ E, swampy meadow, 06.viii.2024, leg. P.V. Golyakov (ALTB; www.inaturalist.org/observations/234189067).

Distribution. The native range of this species is Europe to Central Siberia and Himalaya, Africa, Brazil to north-east Argentina.

Note. This is the first record of the species in the Altai Territory. It is an annual and grows primarily in the temperate biome. In Siberia, the species is very rare, noted only in two habitats in two provinces: Western Siberia Hemiboreal and Altai-Yenisei Mountain-Hemiboreal (Krasnoyarsk Territory). In addition, as an alien species, the plant is noted in Manchurian Continental province (Shekhovtsova 2024).

Family Droseraceae

Drosera rotundifolia L.

Material examined. Russia, Altai Territory, Talmensky District, near the village of Vypolzovo, Lake Klyukvennoye, 53°50ʹ11.6ʺ N, 83°46ʹ46.1ʺ E, cranberry-sphagnum bog, 01.vii.2025, leg. P.V. Golyakov (ALTB).

Distribution. Occurs in most regions of the Holarctic.

Note. This species is rare in the Altai Territory, with only 11 occurrences previously known in the region (Terekhina and Kopytina 2016). The main habitats of Drosera rotundifolia are acidic bogs and poor fens, although it has also been recorded in intermediate-rich and extreme-rich fens. It predominantly grows in Sphagnum-dominated communities (Baranyai and Joosten 2016). Due to the ongoing degradation of sphagnum bogs in the Altai region, D. rotundifolia faces a threat of extinction within the Altai Territory.

Lichens

Family Collemataceae

Lathagrium dichotomum (With.) Otálora, P.M. Jørg. & Wedin

Material examined. Russia, Altai Territory, Zmeinogorsky District, Tigirek Strict Reserve, Tigirek Range, Belaya River, near Krokhalikha River mouth, on boulder in the water, leg. E.A. Davydov (19442) and Y.V. Storozhenko (TIGZ).

Distribution. This species exhibits a circumboreal distribution, occurring across Europe, Asia, and North America. Within Russia, it is documented in the Urals (Paukov and Teptina 2012), Eastern and Southern Siberia (Zhurbenko 2000; Makryi 2008; Vershinina et al. 2012; Urbanavicius and Stepanov 2022), and the northern European part, including Karelia (Thüs and Schultz 2009).

Notes. During a survey of approximately 22 km of the Belaya River channel between the tributaries Kazach'ya Slesarka and Strizhanka, only one population of L. dichotomum was observed near the mouth of the Krokhalikha River. The area occupied by this population was approximately 25 dm², estimated to comprise around 250 individuals.

Lathagrium dichotomum has been recently included in the Red Data Book of the Russian Federation (2024); the report for the Altai Territory was based on unpublished data by Yakovchenko and lacked adequate scientific documentation. Here we amend this inaccuracy by providing detailed evidence of L. dichotomum as a new and confirmed record for the Altai Territory.

This is a widespread but infrequently recorded freshwater species. It thrives on permanently inundated siliceous boulders in clear, cold mountain streams, rivers, large creeks, and lakes with relatively slow-moving water (Nimis et al. 2018). It can also rarely occur in the splash zone of watercourses that experience very infrequent drying (Thüs and Schultz 2009).

Family Coniocybaceae

Chaenotheca chlorella (Ach.) Müll. Arg.

Material examined. Russia, Altai Territory, Eltsovsky District, Salair Ridge, at 6.5 km N from the Kaltyk settlement, at the vicinity of Ivanovka former settlement, 53°17′04″ N, 86°27′44.2″ E, elev. 341 m, 17.vi.2019, swampy spruce (Picea obovata Ledeb.) forest with Carex sp., on wood, leg. E.A. Davydov (22210) and L.S. Yakovchenko (TIGZ).

Distribution. Widely distributed in cool temperate and temperate areas of both the Northern and Southern Hemispheres. Within Russia, this species has been recorded from the Northern and Central European parts, the Northern Urals, and Western Siberia (Urbanavichus 2010; Urbanavichene and Urbanavichus 2019).

Notes. New record for the South Siberia.

Family Lecanoraceae

Myrionora albidula (Willey) R.C. Harris

Material examined. Russia, Altai Territory, Zarinsky District, Salair Ridge, at 13 km to NE from the Novoiushino, Togul River bassin, 53°41′50.6″ N, 85°59′29.3″ E, elev. 317 m, linden (Tilia sibirica Bayer) forest, on bark the fallen Tilia sibirica tree, 21.viii.2020, leg. E.A. Davydov (21413) and Yu.V. Storozhenko (TIGZ).

Distribution. This species inhabits the deciduous and coniferous forest belts of the northern temperate zone. In Russia, it was previously known from the European part (Kostroma, Leningrad, and Tver Regions), the Urals, Siberia (Altai and Trans-Baikal Territoried), and the southern Russian Far East (Palice et al. 2013; Urbanavichene and Urbanavichus 2019; Kotkova et al. 2023). It has been published under the name Biatora albidula Willey for the Caucasus (Urbanavichus et al. 2021). Within the Altai Territory, it was found in Soloneshensky District (Palice et al. 2013).

Notes. New record for the Salair Ridge.

Scoliciosporum perpusillum L.Lahm ex Körb.

Material examined. Russia, Altai Territory, Eltsovsky District, Salair Ridge, at 6.5 km N from the Kaltyk settlement, at the vicinity of former Ivanovka settlement, 53°17′04″ N, 86°27′44.2″ E, elev. 341 m, swampy spruce (Picea obovata) forest with Carex sp., on bark Padus avium Mill., 17.vi.2019, leg. E.A. Davydov (21529) and L.S. Yakovchenko (TIGZ).

Distribution. Within Russia, it is known from several regions in the southern, northern, and central European parts. In Siberia, it has been recorded from the Republic of Altai and Buryatia (Kharpukhaeva and Urbanavichus 2015, Urbanavichus and Urbanavichene 2022). In the Altai Territory, it has been reported for the North-Western Altai (Davydov and Printzen 2012). Outside of Russia, this species is recorded from Central and Northern Europe (Dymytrova 2011).

Notes. New record for the Salair Ridge.

Family Peltigeraceae

Peltigera collina (Ach.) Schrad.

Material examined. Russia, Altai Territory, Eltsovsky District, Salair Ridge, 4.2 km N of Benzherep II settlement, left bank of the Chumysh River, 53°18′6″ N, 86°47′52″ E, elev. 235 m, alone Salix sp. near the road, on soil, 15.vi.2019, leg. E.A. Davydov (17998) and L.S. Yakovchenko (TIGZ).

Distribution. This species is mainly distributed in the temperate zone of the northern hemisphere. In Russia: Arctic, European part, Caucasus, Urals, Siberia and Far East. (Urbanavichus 2010). In the Altai Territory the species has been reported for North-Western and Northern Altai (Davydov 2001; Davydov and Konoreva 2015). On the Salair Ridge the species was known from the Novosibirsk Region (Sedelnikova 2007).

Notes. New record for the Salair botanical-georgaphical province of the Altai Territory.

Peltigera extenuata (Nyl. ex Vain.) Lojka

Material examined. Russia, Altai Territory, Zarinsky District, Salair Ridge, ca. 1 km SE of Alambai Settlement, 54°01′38″ N, 85°54′06″ E, elev. 235 m, Abies sibirica – Pinus sibirica forest with Populus tremula and Betula pendula, on rocks, 13.ix.2019, leg. E.A. Davydov (22321) and L.S. Yakovchenko (TIGZ).

Distribution. Cosmopolitan species widely distributed in Russia, where it was recorded in the Caucasus, Central Russia, Siberia, Kamchatka (Urbanavichus 2010; Ismailov 2021; Muchnik and Kazakova 2020). In the Altai Territory was found in North-West Altai (Davydov 2001).

Notes. New record for the Salair Ridge.

Family Physciaceae

Physcia alnophila (Vain.) Loht. et al.

Material examined. Russia, Altai Territory, Eltsovsky District, Salair Ridge, at 6.5 km N from the Kaltyk settlement, at the vicinity of former Ivanovka settlement, 53°16′50.6″ N, 86°27′53.7″ E, elev. 341 m, riparian willow thickets, on branches of Salix sp., 17.vi.2019, leg. E.A. Davydov (21434) and L.S. Yakovchenko; Zarinsky District, at 13 km to NE from the Novoiushino, Togul River bassin, 53°41′50.9″ N, 85°59′48.5″ E, elev. 310 m, linden (Tilia sibirica) forest, on the bark of Tilia sibirica, 21.viii.2020, leg. E.A. Davydov (21495, 21534) and Yu.V. Storozhenko (TIGZ).

Distribution. The species is widespread, occurring in Europe, Asia, and North America. In Russia, Physcia alnophila is common in the northern European part, the Urals, Siberia, and throughout the Russian Far East (Urbanavichene 2008; Stepanchikova et al. 2010, Galanina et al. 2017). Within the Altai Territory, it has been found in the northwestern part of the region, including Tigirek Strict Reserve (Davydov 2012).

Notes. New record for the Salair Ridge. Specimens of this species are often kept in herbarium under different species names, mainly Physcia aipolia (Galanina et al. 2017).

Physcia tenella (Scop.) DC.

Material examined. Russia, Altai Territory, Zarinsky District, at 13 km to NE from the Novoiushino, Togul River basin, 53°41′58″ N, 85°59′50″ E, elev. 330 m, 11.v.2022, linden (Tilia sibirica) forest, on the bark of of Tilia sibirica, leg. E.A. Davydov and Yu.V. Storozhenko; Togulsky District 3.5 km to NE from the Shumikha, Togul River basin, right bank of the river Togul, between the river Togul and the river Mochishche, 53°38′32.4″ N, 86°01′37.9″ E, elev. 221 m, 18.vi.2019, aspen (Populus tremula L.) forest leg. E.A. Davydov (22183, 21533) and L.S. Yakovchenko (TIGZ).

Distribution. This species is primarily distributed within the temperate zone of the Northern Hemisphere. In Russia, it is found across the Arctic, European part, Caucasus, Urals, Siberia, and the Far East. The species is widespread throughout Russia (Urbanavichus 2010). Within the Altai Territory, the species is reported for the North-Western and Northern Altai regions (Davydov 2001; Davydov and Konoreva 2015). On the Salair Ridge, it is known from the Novosibirsk Region (Sedelnikova 2007).

Notes. New record for the Salair botanical-georgaphical province of the Altai Territory.

Insects, Lepidoptera

Family Tortricidae

Cydia medicaginis (Kuznetzov, 1962)

Material examined. 1 male, Russia, Altai Territory, Krasnoshchekovo District, Chinetinsky Reserve, 14 km SE of village Chineta, Forest-steppe, 775 m, 24.vii.2023, 51.23720° N, 83.17714° E, leg. P. Pavlova (RYB).

Distribution. Euro-Siberian species (Anikin et al. 2019).

Notes. New to Altai Territory.

Cochylis pallidana Zeller, 1847

Material examined. 1 male, Russia, Altai Territory, Charysh District, 8 km S of village Pokrovka, lake Ozernoe, 1500 m, 17–18.vii.2025, 51.045834° N, 83.645250° E, leg. P. Pavlova (RYB).

Distribution. Trans-Palearctic species (Anikin et al. 2019).

Notes. New to Altai Territory.

Aethes cnicana (Westwood, 1854)

Material examined. 1 male, Russia, Altai Territory, Salair Mts., Togul District, 31 km NE of village Togul, linden (Tilia sibirica) forest, 370 m, 17.ix.2023, 53.69726° N, 85.99494° E, leg. P. Pavlova (RYB).

Distribution. Trans-Palearctic species (Anikin et al. 2019).

Notes. New to Altai Territory.

Family Chimabachidae

Dasystoma salicella (Hübner, 1796)

Material examined. 2 males, Altai Territory, Tal'menka district, Tal'menka village vicinity, Chumysh river valley, 53.795159 N, 83.544224 S, 15.iv.2025, S.A. Knyazev (CSKO).

Distribution. From Europe to Far East (Anikin et al. 2019). In Siberia known from Omsk Province, Republic of Altai (Knyazev 2022; Knyazev and Ivonin 2025) and Altai Territory.

Notes. New to Altai Territory. Males were attracted at pheromones.

Family Cossidae

Dyspessa salicicola (Eversmann, 1848)

Material examined. 2 males, Russia, Altai Territory, near Barnaul, South-Siberian Botanical Garden, 53.263832° N, 83.671093° E, 18.vi.2025, leg. A. Slepchenko (RYB).

Distribution. Nominative subspecies distributed in Bulgaria, Macedonia, Albania, Greece, Ukraine, SW Russia, Transcaucasia, Turkey, Сentral Kazakhstan (Romanoff 1885; Spuler 1910; Silbernagel 1944; Daniel 1962; Didmanidze 1978; Didmanidze and Zurashvili, 1981; de Freina 1983, 1996; Efetov and Budashkin 1990; de Freina and Witt, 1990; Yakovlev 2004, 2005, 2009, 2011; Didmanidze and Yakovlev, 2007; Kazenas and Baizanov 2009; Bayanov et al. 2015; Yakovlev et al. 2015, 2020, 2022; Yakovlev and Witt, 2016; Streltzov et al. 2022).

Notes. New species for Western Siberia. Possibly introduced accidentally.

Family Erebidae

Catocala helena Eversmann, 1856

Material examined. 15 specimens, Russia, Altai Territory, near Novoaltaisk, 53.444054° N, 83.939779° E, 2–13.viii.2024, 27.vii–27.viii.2025, leg. A. Slepchenko (RYB).

Distribution. Manchurian nemoral-subboreal species historically distributed in East Palaearctic (China, Korea, Mongolia, south of the Russian Far East and Transbaikalia) (Kononenko 2010). The species was recently found in West Siberia, South Ural and the south of the European part of Russia and Kazakhstan (Karaganda and Pavlodar Provinces) (Knyazev 2011; Ismagilov and Krivosheev 2020; Knyazev et al. 2021; Titov et al. 2022).

Notes. New species for Altai Territory.

Catocala bella Butler, 1877

Material examined. 1 male, Russia, Altai Territory, near Novoaltaisk, 53.444054° N, 83.939779° E, 12.viii.2025, leg. A. Slepchenko (RYB).

Distribution. Manchurian nemoral-subboreal species historically distributed in East Palaearctic (China, Korea, Mongolia, south of the Russian Far East and Transbaikalia) (Kononenko 2010). The species was recently found in West Siberian Plain (Knyazev 2020; Knyazev et al. 2022).

Notes. New species for Altai Territory.

Hypena crassalis (Fabricius, 1787)

Material examined. 1 male, Russia, Altai Territory, near Novoaltaisk, 53.444054° N, 83.939779° E, 13.vi.2025, leg. A. Slepchenko (RYB).

Distribution. European species. In Siberia known from Baikal Region and Altai Territory (Matov and Belova 2016; Gordeev et al. 2022)

Notes. New species for Altai Territory.

Family Noctuidae

Conistra rubiginea (Denis & Schiffermüller, 1775)

Material examined. 19 specimens, Russia, Altai Territory, Barnaul, 53°15′07.0″ N, 83°44′22.5″ E, 6–7.v.2022, 28.iv–5.v.2024, 26.iv.2025, leg. T. Zalutsky (TZB); 1 specimen, Altai Territory, Barnaul, 53°12′09.4″ N, 83°41′32.0″ E, 2.v.2022, leg. O. Kudrov (OKB); 22 specimens, Russia, Altai Territory, Rebrikha village, 28–30.iv.2024, 53°05′14.9″ N 82°22′37.4″ E, leg. E. Svirin (ESB).

Distribution. European species. From Asia was previously known from Kurgan (Zolotarenko and Dubatolov 2000) and Omsk (Knyazev 2020) regions of West Siberia, also in Pavlodar Region in Kazakhstan (Titov et al. 2017).

Notes. New species for Altai Territory.

Acronicta major (Bremer 1861)

Material examined. 2 specimens, Russia, Altai Territory, near Novoaltaisk, 53.444054° N, 83.939779° E, 7.vi.2025, 4.vii.2025, leg. A. Slepchenko (RYB); 1 male, Altai Territory, near Barnaul, South-Siberian Botanical Garden, 53.263832°N, 83.671093°E, 18.vi.2025, leg. A. Slepchenko (RYB)

Notes. New localities for protected species.

Orthosia cerasi (Fabricius, 1775)

Material examined. 2 male, Russia, Republic of Altai, Elekmonar village, 51°27′10.8″ N, 86°02′39.4″ E, 30.v.2025, leg. T. Zalutsky (TZB).

Distribution. Euro-Siberia species (Anikin et al. 2019).

Notes. New species for Republic of Altai.

Family Lasiocampidae

Dendrolimus pini (Linnaeus, 1758)

Material examined. 2 females, Russia, Altai Territory, Tal'menka district, Ozerki village, 53°38′52.6″ N, 83°37′15.6″ E, 17–18.vi.2025, leg. O. Kudrov (OKB).

Distribution. Euro-Siberian species. From Siberia was previously known from Krasnoyarsk and Baikal Regions (Zolotuhin 2015; Anikin et al. 2019).

Notes. New species for Altai Territory.

Family Sphingidae

Agrius convolvuli (Linnaeus, 1758)

Material examined. 2 male, Russia, Altai Territory, Belokurikha, 52°00′00.6″N, 85°01′13.6″ E, 13.viii.2024, leg. T. Zalutsky (TZB).

Distribution. Cosmopolitan species. Eurasia, Africa, Australia and Oceania (Pittaway 2020; Pittaway and Kitching 2020). Though in Siberia and the adjacent countries it was recorded only locally. Thus, for Mongolia, the species is known on the unique specimen collected in the Trans-Altai Gobi (Yakovlev et al. 2015). It is regularly found in Omsk region (Knyazev 2020). The first find in Altai Territory was published in Yakovlev and Volgin 2020.

Notes. Second record for Altai Territory.

Macroglossum stellatarum (Linnaeus, 1758)

Material examined. 2 male, Russia, Republic of Altai, Uznenya village, 51°31′50.7″ N, 85°56′01.6″ E, 31.v.2025, leg. T. Zalutsky (TZB).

Distribution. Transpalearctic species, a noted summer migrant to the north (Yakovlev et al. 2015; Anikin et al. 2019; Pittaway and Kitching 2023).

Notes. The spring discovery may indicate the formation of stable populations in the Republic of Altai.

The studies of E.A. Davydov, Y.V. Storozhenko and L.S. Yakovchenko were conducted within the framework of the grant of the Russian Science Foundation No. 25-24-20159 and Altai Territory.

• Anikin VV, Baryshnikova SV, Beljaev EA, Budashkin YuI, Van Nieukerken EI, Dubatolov VV, Efetov KA, Zolotuhin VV, Knyazev SA, Kovtunovich VN, Kozlov MV, Kononenko VS, Lovtsova JuA, Lukhtanov VA, Lvovsky AL, Matov AYu, Mironov VG, Nedoshivina, S.V., Ponomarenko MG, Sviridov VA, Sinev SYu, Solovjev AV, Streltsov AN, Trofimova TA, Ustjuzhanin PY, Shovkoon DF, Yakovlev RV (2019) Catalogue of the Lepidoptera of Russia. St. Petersburg, 448 pp. [In Russian]
• Artemov IA (2018) Dichostylis micheliana (L.) Nees (1834). In: Red Book of the Novosibirsk Region: Animals, Plants and Fungi. Andrey Khristolyubov Printing House, Novosibirsk, 376 p. [In Russian]
• Baranyai B, Joosten H (2016) Biology, ecology, use, conservation and cultivation of round-leaved sundew (Drosera rotundifolia L.): a review. Mires and Peat 18: 1–28. https://doi.org/10.19189/MAP.2015.OMB.212
• Bayanov MG, Kniss VA, Khabibulin VF (2015) Catalogue of Animals of Bashkortostan. Ufa, 348 pp. [In Russian]
• Daniel F (1962) Monographie der palaearktischen Cossidae. VI. Genus Dyspessa Hbn. Erster Teil. Mitteilungen der Münchner Entomologischen Gesellschaft 52: 1–38.
• Didmanidze EA (1978) Lepidoptera of arid landscapes of Georgia (Heterocera). Tbilisi, 318 pp. [In Russian]
• Didmanidze EA, Yakovlev RV (2007) Cossidae (Lepidoptera) of Georgia. Entomofauna 28(1): 1–16.
• Didmanidze EA, Zurashvili TM (1981) Materials on study of Macrolepidoptera of Vashlovanskii Reserve. Zapovedniki Gruzii 5: 76–118. [In Russian]
• Davydov EA (2001) An annotated list of lichens of the western part of Altai (Russia). Novosti sistematiki nizshikh rastenii [Novitates Systematicae Plantarum non Vascularium] 35: 140–161. [In Russian]
• Davydov EA (2012) Lichens of the Tigirek Nature Reserve. Proceedings of the Tigirek Nature Reserve 4: 72–89. https://doi.org/10.53005/20767390_2011_4_72 [In Russian]
• Davydov EA, Konoreva LA (2015) Lichens of North Altai Province (Altaisky krai). Vestnik Altaiskoy nauki 1: 197–201. [In Russian]
• Davydov EA, Kosachev P, Golyakov P, Zalutsky T, Svirin E, Kudrov O, Pavlova P, Storozhenko Y, Yakovchenko L, Yakovlev R (2023) New and noteworthy records of Plants, Lichens and Lepidoptera in Altai Territory and Republic of Altai (Southern Siberia). Acta Biologica Sibirica 9: 243–264. https://doi.org/10.5281/zenodo.7865738
• Davydov EA, Printzen Ch (2012) Additions to the lichen biota of the Altai Mountains (Siberia). III. Turczaninowia 15 (1): 85–91.
• Davydov EA, Smirnova LY, Storozhenko YV, Zyatnina MV, Ryzhkova PY, Yakovchenko LS (2022) New localities of protected lichen species on the Salair Ridge in Altai Territory. Acta Biologica Sibirica 8: 143−153. https://doi.org/10.5281/zenodo.7700540
• Dymytrova LV (2011) Notes on the genus Scoliciosporum(Lecanorales, Ascomycota) in Ukraine. Polish Botanical Journal 56(1): 61–75.
• Efetov KA, Budashkin YI (1990) Lepidoptera of Crimea (Macroheterocera). Tavria, Simferopol, 112 pp. [In Russian]
• Freina JJ de (1983) 4. Beitrag zur systematischen Erfassung der Bombyces- und SphingesFauna Kleinasiens. Neue Kenntnisse über Artenspektrum, Systematik und Nomeklatur sowie Beschreibungen neuer Taxa. Mitteilungen der Münchner Entomologischen Gesellschaft 72: 57–127.
• Freina JJ de (1996) Cossidae. In: Karsholt O, Razowski J (Eds) The Lepidoptera of Europe. A distributional Checklist. Apollo Books, Soro, 129–130.
• Freina JJ de, Witt TJ (1990) Die Bombyces und Sphinges der Westpalaearktis (Insecta, Lepidoptera). Bd. 2. Cossoidea, Hepialoidea, Pyraloidea, Zygaenoidea. Edition Forschung & Wissenschaft, München, 134 p.
• Galanina I, Ezhkin A, Yakovchenko L, Himelbrant D, Zheludeva E, Skirina I (2017) Physcia alnophila (Vain.) Loht. et al. in the Russian Far East. Turczaninowia 20(1): 99−106. https://doi.org/10.14258/turczaninowia.20.1.8
• Gerasimovicz LV (2012) Scheuchzeriaceae. Key to plants of the Altai Territory. Publishing House of the Siberian Branch of the Russian Academy of Sciences, Novosibirsk, 483 p. [In Russian]
• Gordeev SY, Gordeeva TV, Rudykh SG (2022) Species Composition of Lepidoptera from Erebidae, Nolidae, and Noctuidae Families (Noctuoidea, Lepidoptera) in Western Transbaikalia (the Republic of Buryatia). Nature of Inner Asia 1(20): 7–71. https://doi.org/10.18101/2542-0623-2022-1-7-71 [In Russian]
• Ismagilov NN, Krivosheev MM (2020) Catocala helena Eversmann, 1856 (Lepidoptera: Erebidae) – a new acclimatized species in the South Ural fauna. Rare and endangered species of animals and plants of the republic of Bashkortostan 30: 20–22. [In Russian]
• Ismailov AB (2021) New species of lichens for Dagestan. Botanicheskiy Zhurnal 106(1): 77−80. [In Russian]
• Kazenas VL, Bayzhanov MKh (2009) Insects of Korgalzhyn Reserve and adjacent territories. Almaty, 270 pp. [In Russian]
• Kharpukhaeva TM (2017) Epiphytic lichens inhabiting Chosenia arbutifolia in the Republic of Buryatia. Priroda Vnutrennei Azii 1(2): 37–42. [In Russian]
• Kharpukhaeva TM, Urbanavichyus GP (2015) Findings of new and rare species of lichens for the Republic of Buryatia. Botanicheskiy Zhurnal 100(8): 850–854. [In Russian]
• Knyazev SA (2011) The first record of Catocala helena Eversmann, 1856 (Lepidoptera: Noctuidae) in Cisbaikalia. Eversmannia 27(8): 126. [In Russian]
• Knyazev SA (2020) Catalogue of Lepidoptera of Omsk Oblast (Russia). Macrolepidoptera. Families: Hepialidae, Brachodidae, Cossidae, Sesiidae, Limacodidae, Zygaenidae, Thyrididae, Drepanidae, Uraniidae, Geometridae, Lasiocampidae, Lemoniidae, Endromididae, Saturniidae, Sphingidae, Notodontidae, Lymantriidae, Arctiidae, Syntomidae, Erebidae, Nolidae, Noctuidae, Hesperiidae, Papilionidae, Pieridae, Lycaenidae, Nymphalidae, Satyridae. Acta Biologica Sibirica 6: 139–226. https://doi.org/10.3897/abs.6.e53005
• Knyazev SA (2022) Catalogue of Lepidoptera of Omsk Region (Russia). Microlepidoptera. Families: Eriocraniidae, Nepticulidae, Opostegidae, Adelidae, Prodoxidae, Incurvariidae, Psychidae, Tineidae, Roeslerstammiidae, Bucculatricidae, Gracillariidae, Yponomeutidae, Argyresthiidae, Plutelliidae, Acrolepiidae, Glyphipterigidae, Ypsolophidae, Lyonetiidae, Bedelliidae, Ethmiidae, Depressariidae, Elachistidae, Parametriotidae, Scythrididae, Chimabachidae, Cryptolechiidae, Oecophoridae, Batrachedridae, Coleophoridae, Momphidae, Blastobasidae, Autostichidae, Cosmopterigidae, Gelechiidae, Pterophoridae, Epermeniidae, Choreutidae, Galacticidae, Tortricidae, Pyralidae, Crambidae. Acta Biologica Sibirica 8: 17–87. https://doi.org/10.5281/zenodo.7690616
• Knyazev SA, Gorbunov PY, Melyakh SF, Nedoshivina SV, Grebennikov ND, Matov AY (2021) The first record of Catocala helena Eversmann, 1856 (Lepidoptera, Erebidae) in Europe with notes on its distribution in Russia. Ecologica Montenegrina 40: 68–74. https://doi.org/10.37828/em.2021.40.5
• Knyazev SA, Ivonin VV (2025) New and interesting records of early spring and late autumn Heterocera species (Insecta, Lepidoptera) in Altai Republic (Russia, South Siberia). Acta Biologica Sibirica 11: 1077–1090. https://doi.org/10.5281/zenodo.17396816
• Knyazev SA, Ivonin VV, Saykina SM (2022) New records of rare and local butterflies and moths (Insecta, Lepidoptera) in Omsk and Novosibirsk regions. Amurian Zoological Journal 14(2): 321–334. https://doi.org/10.33910/2686-9519-2022-14-2-321-334 [In Russian]
• Kononenko VS (2010) Noctuidae Sibiricae. Vol. 2. Micronoctuidae, Noctuidae: Rivulinae – Agaristinae. Entomological Press, Sorø, 475 pp.
• Konoreva L, Chesnokov S, Yakovchenko L, Ohmura Y, Davydov E (2020) New records to the lichen biota of Russia, 1 – Sakhalin Region, with new records for the Russian Far East and the Asian part of Russia. Botanica Pacifica 9(2): 161–173. https://doi.org/10.17581/bp.2020.09203
• Kosachev PA, Albach DC (2022) Floristic novelties of Veronica subgenus Pseudolysim achium in Altai. Turczaninowia 25(3): 153–158. https://doi.org/10.14258/turczaninowia.25.3.14
• Kotkova VM, Czernyadjeva IV, Davydov EA, Doroshina GY, Efimov DY, Efimova LA, Frolov IV, Gabiger YI, Glushсhenko MY, Gorbunova IA, Himelbrant DE, Ignatenko ME, Kalinina LB, Kurbatova LE, Kushnevskaya HV, Lashchinsky NN, Lotiev KY, Moroz EL, Notov AA, Novozhilov YK, Otmakhov YS, Plikina NV, Popova NN, Potemkin AD, Putilina VA, Ryzhkova PY, Sambyla CN, Smirnova EV, Stepanchikova IS, Storozhenko YV, Troeva EI, Tsurykau AG, Vishnyakov VS, Vlasenko AV, Vlasenko VA, Volkova EA, Volosnova LF, Yakovchenko LS, Yatsenko-Stepanova TN, Zhuykov KA, Zueva AS (2023) New cryptogamic records. 11. Novosti sistematiki nizshikh rastenii 57(1): 155–204. https://doi.org/10.31111/nsnr/2023.57.1.155
• Krasnoborov IM, Korotkova EI (1988) Scheuchzeriaceae. Flora Sibiriae. Lycopodiaceae – Hydrocharitaceae. Vol. 1. Nauka, Novosibirsk, 111 p. [In Russian]
• Makryi TV (2008) Lichens. In: Cryptogams of the Baikal National Park. "Geo"Publishing House, Novosibirsk, 113–259.
• Matov AY, Belova NA (2016) To the fauna and ecology of Bombycoidea and Noctuoidea Moths (Lepidoptera: Lasiocampidae, Endromididae, Sphingidae, Notodontidae, Erebidae, Nolidae, Noctuidae). Amurian Zoological Journal 8(1): 52–63. https://doi.org/10.33910/1999-4079-2016-8-1-52-63 [In Russian]
• Muchnik EE, Kazakova MV (2020) New finds of rare and protected lichens in the Ryazan region (Russia). Bulletin of Perm University. Series Biology 4: 264–271. https://doi.org/10.17072/1994-9952-2020-4-264-271
• Palice Z, Printzen Ch, Spribille T, Svensson M, Tønsberg T, Urbanavichene I, Yakovchenko LS, Ekman S (2013) Taxonomy of the genus Myrionora, with a second species from the Southern Hemisphere. Lichenologist 45(2): 159–167. https://doi.org/10.1017/S0024282912000692
• Paukov A, Teptina A (2012) New records of lichens from Middle Urals, Russia. Folia Cryptogamica Estonica 49: 39–43.
• Pittaway AR (2020) Sphingidae of Western Palearctic. http://tpittaway.tripod.com/sphinx/list.htm
• Pittaway AR, Kitching IJ (2020) Sphingidae of the Eastern Palaearctic (including Siberia, the Russian Far East, Mongolia, China, Taiwan, the Korean Peninsula and Japan). http://tpittaway.tripod.com/china/china.htm
• Red Book of the Tambov Region (2019) Mosses, vascular plants, fungi, lichens. Second Edition. ООО "ТPS", Tambov, 480 pp. [In Russian]
• Romanoff NM (1885) Les Lepidopteres de la Transcaucasie. Deuxième partie. Mémoires sur les Lépidoptères par N. M. Romanoff 2: 1–118.
• Sedelnikova NV (2007) Flora of the Salair Ridge. Lichens. "Geo" Publishing House, Novosibirsk, 98–136. [In Russian]
• Seregin AP (Ed.) (2024) Digital herbarium of Moscow State University: Electronic resource. Access mode: https://doi.org/10.17581/bp.plant.depo.msu.ru/ (access date 05/04/2024.13S01)
• Silantieva MM (2013) A checklist of the flora of the Altai region: monograph. Altai Publishing House, Barnaul, 520 pp. [In Russian]
• Silbernagel A (1944) Die Schmetterlinge der Ochrid-Gegend in Macedonien. Zeitschrift der Wiener Entomologischen Gesellschaft 29: 184–187.
• Shekhovtsova IN (2024) Cyperaceae. Checklist of vascular plants of Asien Russia. Botanica pacifica. A journal of plants science and conservation 13 (Special issue): 3–310.
• Shmakov AI (2005) Cystopteridaceae. Flora Altaica. Vol. 1. AzBuka, Barnaul, 205–220. [In Russian]
• Spuler A (1910) Die Schmetterlinge Europas, 2. E. Schweizerbart, Stuttgart, 523 pp.
• Stepanchikova IS, Himelbrant DE, Kukwa MV, Kuznetsova ES (2010) Additions to the lichen flora of protected areas Gulf of Finland (within St. Petersburg). Novosti sistematiki nizshikh rasteniy [Novitates Systematicae Plantarum non Vascularium] 44: 237–244. https://doi.org/10.31111/nsnr/2010.44.237 [In Russian]
• Streltzov AN, Ustjuzhanin PYa, Morozov PS, Naydenov AE, Spitsyn VM, Yakovlev RV (2022) Lepidoptera of South Ossetia (Northern Transcaucasia). Part II. Cossidae, Limacodidae, Erebidae (Lymantriinae, Arctiinae, Syntominae, Notodontinae), Lasiocampidae, Lemoniidae, Saturniidae, Sphingidae, Drepanidae and Cimeliidae. Acta Biologica Sibirica 8: 647–654. https://doi.org/10.14258/abs.v8.e40
• Terekhina TA, Kopytina TM (2016) Drosera rotundifolia L. In: Red Data Book of the Altai Territory. Vol. 1. Rare and endangered plant and fungi species. Altai State University Publishing, Barnaul, 101 p. [In Russian]
• Timokhina SA, Bondareva NV (1990) Rhynchospora Vahl – Ocheretnik. Flora Sibiriae. Cyperaceae. Vol. 3. Nauka, Novosibirsk, 31 p. [In Russian]
• Titov SV, Volynkin AV, Dubatolov VV, Černila M, Reznichenko SM, Bychkov VS (2017) Noctuoid moths (Lepidoptera: Erebidae, Nolidae, Noctuidae) of North-East Kazakhstan (Pavlodar Region). Ukrainian Journal of Ecology 7(2): 142–164. https://doi.org/10.15421/2017_32
• Titov SV, Volynkin AV, Pulikova GI, Kaptyonkina AG (2022). The first record of Catocala helena Eversmann, 1856 from Kazakhstan (Lepidoptera: Noctuoidea: Erebidae: Catocalini). Ecologica Montenegrina 52: 42–48. https://doi.org/10.37828/em.2022.52.6
• Urbanavichene I, Urbanavichus G (2019) New records of lichens and allied fungi from the Kostroma Region, Russia. Folia Cryptogamica Estonica 56: 53–62. https://doi.org/10.12697/fce.2019.56.06
• Urbanavichene IN (2008) Genus Physcia(Schreb.) Michx. In: Opredelitel lishaynikov Rossii [Handbook of lichen of Russia]. Nauka, St. Petersburg 10: 258–276. [In Russian]
• Urbanavichus G (2010) A checklist of the lichen flora of Russia. Nauka, St. Petersburg, 194 pp. [In Russian]
• Urbanavicius GP, Stepanov NV (2022) Collema dichotomum. In: Red Book of the Krasnoyarsk Territory. In 2 vols. Vol. 2: Rare and endangered species of wild plants and fungi. Siberian Federal University, Krasnoyarsk, 588 p. [In Russian]
• Urbanavichus GP, Urbanavichene IN (2022) The core of the Kologriv Forest State Nature Reserve (Russia) is a hotspot of lichen biodiversity in the southern taiga of Eastern Europe. Nature Conservation Research 7(3): 46–63. https://doi.org/10.24189/ncr.2022.029 [In Russian]
• Urbanavichus GP, Urbanavichene IN, Vondrák J, Ismailov A (2021) Epiphytic lichen biota of Prielbrusie national park (Northern Caucasus, Russia). Nature Conservation Research 6: 77–94. https://doi.org/10.24189/ncr.2021.048
• Verkhozina AV, Biryukov RYu, Bogdanova ES, Bondareva VV, Chernykh DV, Dorofeev NV, Dorofeyev VI, Ebel AL, Efimov PG, Efremov AN, Erst AS, Fateryga AV, Gamova NS, Glazunov VA, Gudkova PD, Juramurodov IJ, Kapitonova OА, Kechaykin AA, Khapugin AA, Kosachev PA, Krupkina LI, Kulagina MA, Kuzmin IV, Lian Lian, Koychubekova GA, Lazkov GA, Luferov AN, Mochalova OA, Murtazaliev RA, Nesterov VN, Nikolaenko SA, Novikova LA, Ovchinnikova SV, Plikina NV, Saksonov SV, Senator SA, Silaeva TB, Suleymanova GF, Hang Sun, Tarasov DV, Tojibaev KSh, Vasjukov VM, Wei Wang, Zibzeev EG, Zolotov DV, Zykova EYu, Krivenko DA (2021) Findings to the flora of Russia and adjacent countries: New national and regional vascular plant records. Botanica Pacifica 3 10 (1): 1–24. https://doi.org/10.17581/bp.2021.10110
• Vershinina SE, Himelbrant DE, Kuznetsova ES, Gabysheva LM, Gabyshev EM (2012) The first data on the lichen flora of the Olekminsky State Reserve (Republic of Sakha-Yakutia). Bulletin of Tver State University. Series: Biology and Ecology 25(3): 13–149.
• Yakovlev RV (2004) Carpenter-Moths (Lepidoptera, Cossidae) of Siberia. Euroasian entomological journal 3(2): 155–163. [In Russian]
• Yakovlev RV (2005) New data on distribution and systematic of Cossidae (Lepidoptera) of Europe and adjacent territories. Eversmannia 3–4: 18–27. [In Russian]
• Yakovlev RV (2011) Catalogue of the Family Cossidae of the Old World. Neue Entomologische Nachrichten 66: 1–129.
• Yakovlev RV, Gus'kova EV, Doroshkin VV, Titov SV (2015) Sphingidae of the Mongolian Altai (Lepidoptera: Sphingidae). SHILAP Revista de lepidopterologia 43(171): 467–478.
• Yakovlev RV, Poltavsky AN, Ilyina EV, Shchurov VI, Witt Th (2015) Cossidae (Lepidoptera) of the Russian Caucasus with the description of a new species. Zootaxa 4044(2): 270–288. https://doi.org/10.11646/zootaxa.4044.2.5
• Yakovlev RV, Shapoval NA, Ivonin VV, Knyazev SA, Kuftina GN, Masharskiy AE (2020) A new species of Carpenter Moths (Lepidoptera, Cossidae) from Tarbagatai (NE Kazakhstan) and Altai (SW Siberia, Russia) Mountains. Zootaxa 4896(1): 085–095. https://doi.org/10.11646/zootaxa.4896.1.3
• Yakovlev RV, Shapoval NA, Shapoval GN, Naydenov AE (2022) Review of the Dyspessa salicicola (Eversmann, 1848) species group (Lepidoptera, Cossidae: Cossinae). Acta Biologica Sibirica 8: 693–712. https://doi.org/10.14258/abs.v8.e43
• Yakovlev RV, Volgin IG (2020) New finding of Convolvulus hawkmoth – Agrius convolvuli (Linnaeus, 1758) (Lepidoptera, Sphingidae) in the south of Western Siberia. Ecologica Montenegrina 38: 155−157. https://doi.org/10.37828/em.2020.38.23
• Yakovlev RV, Witt TJ (2016) Carpenter-moths of Turkmenistan (Lepidoptera: Cossidae). Biological Bulletin of Bogdan Chmelnitskiy Melitopol State Pedagogical University 6(3): 164–173. https://doi.org/10.15421/201684
• Zhdanov IS (2013) Additions to the lichen flora of Central Siberian Biosphere Reserve (Krasnoyarsk Territory). Novosti sistematiki nizshikh rastenii 47: 200–214. https://doi.org/10.31111/nsnr/2013.47.200 [In Russian]
• Zhurbenko MP (2000) Lichens and lichenicolous fungi of the Putoransky Reserve. Flora and fauna of nature reserves 89: 55. [In Russian]
• Zolotarenko GS, Dubatolov VV (2000) A check-list of Noctuidae (Lepidoptera) of the Russian Part of the West-Siberian Plain. Far Eastern Entomologist 94: 1–23.
• Zolotuhin VV (2015) Lappet Moths (Lepidoptera: Lasiocampidae) of Russia and Adjacent Territories. Korporaciya tekhnologiy prodvizheniya, Ulyanovsk, 384 pp. [In Russian]