Introduction
This work continues a series of the authors' investigations of the Lepidoptera (mainly Sesiidae and Zygaenidae) of the world fauna (see e.g. Efetov 1996a, 1996b, 1996c, 1997, 1998a, 1998b, 1999, 2001a, 2001b, 2006, 2010, 2012, 2018; Efetov et al. 2004, 2011b, 2015a, 2019a, 2019c, 2020, 2023, 2024; Efetov and Savchuk 2009, 2013; Efetov and Tarmann 1999, 2013, 2014a, 2014b, 2016, 2017, 2020, 2024; Gorbunov 2015, 2016, 2017, 2018a, 2018b, 2018c, 2023, 2024a, 2024b, 2024c, 2024d, 2025a, 2025b, 2025c, 2026a, 2026b, 2026d; Gorbunov and Ivanov 2026; Gorbunov and Koshkin 2026, Mirić et al. 2024; Yardım et al. 2024).
As we have noted on several occasions, in the course of an in-depth study of the clearwing moth fauna of the Crimean Peninsula (Gorbunov and Efetov 1990, 2016, 2018, 2024, 2025a, 2025b, 2025c, 2025d, 2026a; Efetov et al. 2012a, 2012b; Efetov and Gorbunov 2021, 2024), three new species were described (Gorbunov and Efetov 2018; Gorbunov 2019a, 2019b), and Bembecia eversmanni O. Gorbunov, 2024, was recorded for the peninsula for the first time (Gorbunov et al. 2025a; Gorbunov and Efetov 2026c). In addition, we have collected a substantial amount of material from various localities across the peninsula, many of which had not previously been surveyed for the presence of taxa of the family Sesiidae. As noted in our recent publications (Gorbunov and Efetov 2024, 2025a, 2025b, 2025c, 2025d, 2026a), we have analyzed all available collection specimens, literature sources containing reliable species identifications, and data from the "Lepidoptera of Crimea" website (Savchuk 2026), as well as, occasionally, information from the online resource iNaturalist (2026).
As is well known from our publications, we have made active use of artificial sex attractants in our work. It is widely recognized that sex pheromones and sex attractants play an increasingly important role in the study of Lepidoptera: they enable researchers to discover new species, identify populations of pests or species rarely represented in collections, and monitor and control the seasonal flight activity of moths (Efetov et al. 2010, 2011a, 2015b, 2018, 2019b, 2020, 2023, 2025; Efetov and Kucherenko 2020, 2021; Drouet et al. 2021; Efetov and Gorbunov 2024; Gorbunov 2015, 2016, 2017, 2018a, 2018b, 2018c, 2022a, 2022b, 2022c, 2022d, 2023; Gorbunov et al. 2025b; Gorbunov and Efetov 2026b; Razov et al. 2017).
This publication focuses on the morphology and distribution of Synanthedon andrenaeformis (Laspeyres, 1801) on the Crimean Peninsula. It is appropriate to point out here that on the Crimean Peninsula three administrative-territorial units of the Russian Federation, viz. Federal City of Sevastopol', the Republic of Crimea and Genichesk District of the Kherson Region (the northern part of the Arabat Spit), are located. We were able to collect this species only in the first two regions.
Materials and methods
The descriptions of the specimens were made using a Leica® EZ4 stereomicroscope with LED illumination. All images were taken using a Sony® α450 DSLR camera equipped with a Minolta® 50 f/2.8 Macro lens. The figures of the genitalia were taken with a Keyence® BZ-9000 Biorevo Fluorescence Microscope. The processing of all illustrations was finalized using Adobe® Photoshop® 2020 software. Pictures of the specimens are labeled with a number which consists of the name of the family, two consecutive digits and a year (e.g. Sesiidae pictures No 0075-0076–2023). These numbers correspond to those of the illustrated specimens in the archives of the first author. The dissected genitalia are equipped with the corresponding number placed in the microtube. This number as a label (e.g. Genitalia preparation No OG–005-2024) is pinned under the specimen and listed in the first author's archive.
The specimens were attracted to artificial female sex pheromones for sesiid moths produced by PHEROBANK®, Wijk bij Duurstede, the Netherlands.
The names of plants were verified with the WFO (2026).
On the distribution map, the locations of records from literature and Internet resources are indicated by rings, and the locations of our own catches and studied specimens from museums are shown by black dots.
The material studied or mentioned below is kept in the following collections abbreviated in the text as follows:
COGM – collection of the A.N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Moscow, Russia; CKES – collection of Konstantin A. Efetov, Simferopol', Russia; CTGA – collection of Theo C. Garrevoet, Antwerp, Belgium.
Results
Synanthedon andrenaeformis (Laspeyres, 1801) was first collected in the Crimea, in the vicinity of Krasnoles'ye, in the Simferopol' District, in June 2019 by the Crimean enthusiast V.V. Savchuk, using artificial sex attractants (Savchuk and Kajgorodova 2020; Savchuk 2026). A couple of years later, we were fortunate enough to collect a few males near Frontovoye, in the vicinity of Sevastopol', also using artificial sex attractants. Subsequently, we managed to collect several larvae and rear a pair of adult moths (Figs 9, 10), the female of which proved to be the first female specimen recorded for the Crimea. In addition to this, we collected a small series of males. All the males collected were drawn to the artificial sex attractants synthesized for two European species: Synanthedon vespiformis (Linnaeus, 1761) and Synanthedon andrenaeformis (Laspeyres, 1801). The males were attracted exclusively during the morning hours, up until approximately 11:00 AM. We succeeded in finding this species in four habitats across two administrative-territorial units of the Crimean Peninsula, viz. the Federal city of Sevastopol' and the Republic of Crimea.
S. andrenaeformis is currently known from five locations on the Crimean Peninsula, all of which are situated within its mountainous region (Fig. 22).
Taxonomic account
Family Sesiidae Boisduval, 1828
Subfamily Synanthedoninae Le Cerf, 1917
Tribe Synanthedonini Le Cerf, 1917
Genus Synanthedon Hübner, 1819
Synanthedon andrenaeformis (Laspeyres, 1801)
Figs 1–22
"Sesia Andrenaeformis.": Laspeyres 1801: 20. Type locality: "Habitat in Hungaria" [= Hungary]. Type material: destroyed (Špatenka et al. 1999).
= "Sphinx… anthraciniformis." [Esper 1800]: Esper 1789–1804: 29, Tab. XLIV, Cont. 19, Fig. 1, 2. Type locality: "in der Gegend von Ofen" [= Hungary: vicinity of Budapest]. Name rejected by ICZN, Opinion Nr. 1287 (ICZN 1984).
= "Troch[ilium]. Allantiforme.": Newman 1832: 79. Type locality: "Habitat in Anglia". Type material: lost (Špatenka et al. 1999).
= "[Sesia F.]… Andreniformis": Staudinger 1861: 17. Unjustified emendation of Sesia andrenaeformis Laspeyres, 1801.
= "Synanthedon perigordensis sp. n.": Garrevoet and Vanholder 1996: 141, figs 1–5, 6a, 7a. Type locality: "France, Dep. Dordogne, Plaisance/Issigeae". Type material: holotype ♂ (CTGA).
Records for the Crimea. Savchuk and Kajgorodova 2020: 260, fig. 24; Laštůvka and Laštůvka 2025: 30, fig. 25, map 25; Savchuk 2026.
Description. Males (Figs 1–6) and females (Figs 7–8) of this species are virtually indistinguishable in terms of colouration. Females are merely somewhat more robust than males. Both males and females show slight variation in the number of yellow and white scales on the abdomen. The basic sizes of both males and females vary as follows: wing span 16.2–23.5 mm; body length 11.0–14.2 mm; forewing length 7.4–11.0 mm; length of antenna 5.6–8.0 mm.
Head: flagellum of antenna and scapus black with dark blue-violet sheen; frons black with dark bronze-violet sheen; vertex black with dark greenish-violet sheen; labial palpus black with dark blue-violet sheen dorsally and white ventrally; occipital fringe black; neck plate black with bright blue-violet sheen and small white spot laterally.
Thorax: patagium black with bright greenish-violet sheen; tegula, meso- and metathorax black with dark blue-violet sheen; besides this, tegula and mesothorax densely covered with thin, hair-like, black scales; thorax laterally dark grey-brown with bright violet sheen and large yellow spot medially; posteriorly, both metepimeron and metameron dark grey-brown with dark violet sheen, covered with black, long, hair-like scales.
Legs: fore coxa black with bright blue-violet sheen, sometimes with narrow white exterior margin; hind tibia black with bright blue-violet sheen and several yellow scales dorsodistally; spurs white.
Forewing: dorsally costal margin and discal spot black with dark violet sheen; CuA-stem, anal margin, veins within external transparent area and apical area black with bronze-violet sheen; cilia dark-brown to black with bronze sheen; ventrally costal margin lemon-yellow with golden hue; CuA-stem and anal margin dark brown to black with bronze-violet sheen and admixture of several lemon-yellow scales; discal spot and surface between veins R1–R3 black with dark violet sheen and several lemon-yellow scales at base of veins M1–M3; veins within external transparent area black with bronze-violet sheen; apical area black with bright bronze-violet sheen and with narrow lemon-yellow stripe between veins R3–CuA1; transparent areas well-developed: posterior transparent area reaching distal margin of discal spot, external transparent area with parallel margins, divided into five cells between veins R3–CuA1, level to vein M2 about 0.8–1.4 times broader than discal spot and about 0.3–0.4 times as broad as apical area; discal spot with parallel margins; cilia dark-brown to black with bronze sheen.
Hindwing transparent; veins, discal spot and outer margin black with dark bronze-violet sheen; cilia dark-brown to black with bronze sheen, anally white; discal spot small, triangular, reaching base of vein M2; outer margin narrow but slightly gradually broadened to costal margin, about as broad as cilia.
Abdomen black with dark greenish-blue sheen; dorsally tergites 2 and 4 with narrow yellow stripe distally; ventrally sternites 4–6 in male and only 4 in female white; anal tuft black with dark blue-violet sheen and straw-yellow, yellow or yellow-orange medio-distally.
Male genitalia (genitalia preparation No OG–005-2024) (Figs 11–14). Tegumenuncus complex (Fig. 11) relatively broad; scopula androconialis well-developed, about 0.6 times as long as length of tegumenuncus complex; crista gnathi medialis long and broad; crista gnati lateralis subcordiform, short and slightly shorter than crista gnathi medialis (Fig. 11); valva (Fig. 12) trapeziform, crista sacculi oblique, long, about 0.3 as long as length of valva, pocket-shaped, covered with one row of strong flat-topped setae; saccus (Fig. 13) narrow, slightly bilobed basally, about as long as vinculum; phallus (Fig. 14) rather thin, straight, slightly broadened subdistally, about 0.67 times as long as valva; vesica with numerous minute cornuti (Fig. 14).
Female genitalia (genitalia preparation No OG–006-2024) (Fig. 15). Papillae anales relatively large, well-sclerotized, with numerous setae; posterior apophysis about 1.2 times as long as anterior apophysis; tergite 8 relatively broad, well-sclerotized, with sparse setae at distal margin; both lamellae postvaginalis and antevaginalis undeveloped; ostium bursae well-sclerotized, distinctly cup-shaped, opening in proximal part of segment 8; antrum well-sclerotized, very short and thin; ductus bursae thin, membranous, long, about 1.5 times as long as posterior apophysis; corpus bursae elongated-ovoid without signa.
Differential diagnosis. Among the clearwing moths of the Crimean fauna, this species could only be confused with members of the genus Tipulia Králíček et Povolný, 1977, of which only two species have been recorded in the region, viz. T. spuleri (Fuchs, 1908) and T. tipuliformis (Clerck, 1759). S. andrenaeformis differs rather well from both these species by its generally larger size and the colouration of the frons (with bright white stripe laterally in species of the genus Tipulia (see figs 2 and 4 in Gorbunov 2024b), vs. frons completely black with dark bronze-violet sheen in S. andrenaeformis) and the abdomen (dorsally, tergites 2, 4, 6, and 7 in the male, and 2, 4, and 6 in the female, each bear a thin yellow stripe distally; ventrally, the abdomen lacks white scales in the species of the genus Tipulia, vs. dorsally only tergite 2 and 4 with thin yellow stripe and ventrally abdomen with white scales in S. andrenaeformis. Compare Figs 1–8 in this article with figs 1 and 3 in Gorbunov 2024b). In T. tipuliformis and T. spuleri, the anal tuft is black and lacks yellow scales, whereas in S. andrenaeformis, the anal tuft always contains straw-yellow or orange-yellow scales. In addition, these species can be easy distinguished from each other by the conformation of the both male and female genitalia; compare Figs 11–15 in this publication with figs 7–11 in Gorbunov (2024b).
Bionomics. In the Crimea, we identify only Viburnum lantana L. (Viburnaceae) as the host plant for this species, although in Europe, V. opulus L. is also listed (Laštůvka and Laštůvka 2025). The young larva lives beneath the bark, but by the autumn, it bores a tunnel into the center of the trunk or branch, where it overwinters. During its second year, the larva extends the tunnel slightly, to a length of about 6–7 cm. Before its second winter, it prepares an exit hole, which is covered by a small, round lid made of bark (Figs 16–21). The larva pupates within the tunnel inside a very loose silk cocoon. The adult moths emerge and fly in June (Figs 9, 10). Males are actively attracted to artificial sex attractants synthesized for the European Synanthedon vespiformis (Linnaeus, 1761).
Habitat. Broad-leaved forests, where the host plant is present, either along the forest edges or deeper within the woods. They may also inhabit garden plots, where they can be found on cultivated Viburnum lantana or V. opulus shrubs.
Distribution. The species occurs locally but is nonetheless quite widely distributed across Western, Northern, and Southern Europe: from Spain, France, and England in the west to Poland, Ukraine (iNaturalist 2024), and Bulgaria in the east, and from England, Belgium, and northern Germany in the north to Italy and northern Greece in the south; additionally, isolated populations are known to exist in southern Norway and Sweden (Laštůvka and Laštůvka 2025). In Russia, apart from the Crimea, there are only reports of this species from the Samara and Nizhnyy Novgorod Regions, and, based on very old specimens, from the Republic of Tatarstan (Sachkov and Popova 2011; Gorbunov 2019c; Shmelev 2020). In the Crimea, this species is known from five localities across two administrative-territorial units: the Federal city of Sevastopol' and the Republic of Crimea (Fig. 22).
Republic of Crimea: 5 ♂♂, Russia, Crimea, Simferopol' District, Kizilovoye, 44°48.957′N, 34°02.346′E, ca 290 m., 16.VI.2023, O.G. Gorbunov and K.A. Efetov leg. (Sesiidae pictures No 0075-0082–2023) (COGM); 8 ♂♂, same locality, 18.VI.2023, O.G. Gorbunov and K.A. Efetov leg. (Sesiidae pictures No 0061-0064–2023, 0113-0124–2023) (CKES, COGM).
City of Sevastopol': 2 ♂♂, Russia, Crimea, Sevastopol', Frontovoye, 45°40.242′N, 033°43.487′E, ca 115 m., 24.VI.2021, O.G. Gorbunov and K.A. Efetov leg. (Sesiidae pictures Noo 0329-0330–2021, 0335-0336–2021) (COGM); 1 ♂, 1 ♀, Russia, Crimea, Sevastopol', Verkhnesadovoye, 44°40.409′N, 33°41.841′E, ca 97 m., 25.V.2022, ex larvae from trunks of Viburnum lantana L. (Viburnaceae); moths emerged 25.VI (♂) and 26.VI.2022 (♀), O.G. Gorbunov leg. (Sesiidae pictures No 0041-0044–2022) (COGM); 1 ♂, Russia, Crimea, Sevastopol', Frontovoye, 44°39.417′N, 33°43.084′E, ca 144 m., 15.VI.2023, O.G. Gorbunov and K.A. Efetov leg. (Sesiidae pictures No 0089-0090–2023) (Genitalia preparation No OG–005-2024) (COGM).
Conclusion
As a result of the research, S. andrenaeformis was found in five localities across two administrative-territorial units of the Russian Federation on the Crimean Peninsula. This species is recorded for the first time in the Federal City of Sevastopol'. The host plant of the species on the Crimean Peninsula is Viburnum lantana L. (Viburnaceae).
Acknowledgements
We would like to express our heartfelt gratitude to Mr Aleksey N. Zamesov and Mr Andrey A. Ermakov (Moscow, Russia) for their assistance in collecting material of clearwing moths. Many thanks to Prof. Dr Andrey V. Yena (Simferopol, Russia) for his help in determining the host plants of the Crimean Sesiidae. We are also indebted to Mr Vladislav M. Nagaev (Simferopol, Russia) for carefully checking the English in this manuscript.
This study by the first author was supported by the Ministry of Science and Higher Education of the Russian Federation (projects No 1022061500172-3-1.6.19) and the state assignment of Lomonosov Moscow State University.
The study was carried out using the equipment of the Joint Usage Center "Instrumental methods in ecology" of the A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences (Moscow, Russia).
References
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Gorbunov OG (2016) Nokona mahawu sp. n., a new clearwing moth species (Lepidoptera, Sesiidae) from North Sulawesi, Indonesia. Russian Entomological Journal 25(2): 161–165. https://doi.org/10.15298/rusentj.25.2.02
Gorbunov OG (2017) On the taxonomy and morphology of Leuthneria ruficincta (Lepidoptera, Sesiidae). Zootaxa 4244(1): 127–136. https://doi.org/10.11646/zootaxa.4244.1.7
Gorbunov OG (2018a) New data on the clearwing moth fauna of the Altai Mountains, Russia, with the description of two new species (Lepidoptera, Sesiidae). Zootaxa 4425(2): 263–282. https://doi.org/10.11646/zootaxa.4425.2.4
Gorbunov OG (2018b) A new species of the genus Taikona Arita et O. Gorbunov, 2001 from the Malay Peninsula (Lepidoptera, Sesiidae). Russian Entomological Journal 27(3): 293–296. https://doi.org/10.15298/rusentj.27.3.10
Gorbunov OG (2018c) A new genus and species of clearwing moth (Lepidoptera, Sesiidae) from Ethiopia. Zootaxa 4497(4): 492–500. https://doi.org/10.11646/zootaxa.4497.4.2
Gorbunov OG (2019a) Two new species of the genus Chamaesphecia Spuler, 1910 (Lepidoptera: Sesiidae), with remarks on the genus. Russian Entomological Journal 28(4): 437–457. https://doi.org/10.15298/rusentj.28.4.13
Gorbunov OG (2019b) A new species of the genus Negotinthia O. Gorbunov, 2001 (Lepidoptera: Sesiidae) from Crimea, with remarks on the genus. Russian Entomological Journal 28(3): 303–311. https://doi.org/10.15298/rusentj.28.3.09
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Gorbunov OG (2022a) A new species of the genus Tyrictaca Walker, 1862 (Lepidoptera: Sesiidae) from West Malaysia with a catalogue of the genus. Zootaxa 5104(1): 125–136. https://doi.org/10.11646/zootaxa.5104.1.7
Gorbunov OG (2022b) New data on clearwing moths (Lepidoptera: Sesiidae) of Sakhalin island. Far Eastern Entomologist 449: 21–28. https://doi.org/10.25221/fee.449.4
Gorbunov OG (2022c) A new species of the genus Scoliokona Kallies et Arita, 1998 (Lepidoptera: Sesiidae) from Halmahera, Indonesia. Russian Entomological Journal 31(2): 182–187. https://doi.org/10.15298/rusentj.31.2.16
Gorbunov OG (2022d) To the morphology of little-known Pseudosesia lecerfi (Hampson, 1919) (Lepidoptera: Sesiidae). Russian Entomological Journal 31(3): 286–292. https://doi.org/10.15298/rusentj.31.3.10
Gorbunov OG (2023) A new species of the genus Macrotarsipodes Le Cerf, 1916 (Lepidoptera: Sesiidae) from Ethiopia. Invertebrate zoology 20(4): 461–466. https://doi.org/10.15298/invertzool.20.4.11
Gorbunov OG (2024a) To the knowledge of the fauna of Lepidoptera of Mongolia. Family tussock moths (Lepidoptera: Noctuoidea: Lymantriidae). Caucasian Entomological Bulletin 20(1): 47–59. https://doi.org/10.5281/zenodo.10869524 [In Russian]
Gorbunov OG (2024b) On the systematics of the genus Synanthedon Hübner, 1819 sensu lato (Lepidoptera: Sesiidae). Part I. Tipulia Králíčeket Povolný, 1977. Invertebrate Zoology 21(2): 232–238. https://doi.org/10.15298/invertzool.21.2.10
Gorbunov OG (2024c) On the systematics of the genus Synanthedon Hübner, 1819 sensu lato (Lepidoptera, Sesiidae). Part II. Thamnosphecia Spuler, 1910. Acta Biologica Sibirica 10: 1487–1497. https://doi.org/10.5281/zenodo.14292276
Gorbunov OG (2024d) On the systematics of the genus Synanthedon Hübner, 1819 sensu lato (Lepidoptera: Sesiidae). Part III. Conopia Hübner 1819. Invertebrate Zoology 21(4): 502–514. https://doi.org/10.15298/invertzool.21.4.06
Gorbunov OG (2025a) To the knowledge of the fauna of Lepidoptera of Mongolia. The family prominent moths (Lepidoptera: Noctuoidea: Notodontidae) Caucasian Entomological Bulletin 21(1): 41–57. https://doi.org/10.5281/zenodo.15078733 [In Russian]
Gorbunov OG (2025b) A redescription of Bembecia strandi (Lepidoptera: Sesiidae). Zoosystematica Rossica 34(1): 138–148. https://doi.org/10.31610/zsr/2025.34.1.138
Gorbunov OG (2025c) To the knowledge of the fauna of Lepidoptera of Mongolia. The family forester and burnet moths (Lepidoptera: Zygaenidae) Caucasian Entomological Bulletin 21(2): 201–211. https://doi.org/10.5281/zenodo.17200399 [In Russian]
Gorbunov OG (2026a) A new genus and species of clearwing moth of the tribe Paranthrenini (Lepidoptera: Sesiidae) from Southern Thailand. Invertebrate Zoology 23(1): 136–145. https://doi.org/10.15298/invertzool.23.1.08
Gorbunov OG (2026b) To the knowledge of the fauna of Lepidoptera of Mongolia. Family lappet moths (Lasiocampidae) and saturniids (Saturniidae). Caucasian Entomological Bulletin 22(1): 213–231. https://doi.org/10.5281/zenodo.21642943 [In Russian]
Gorbunov OG (2026c) A new species of the genus Chamaesphecia Spuler, 1910 (Lepidoptera: Sesiidae) from Mongolia. Invertebrate Zoology 23(3): 505–513. https://doi.org/10.15298/invertzool.23.3.08
Gorbunov OG (2026d) New data on the Sesiini of the Oriental and Australian regions (Lepidoptera: Sesiidae) with the establishment of a new genus. Russian Entomological Journal 35(3): 408–422. https://doi.org/10.15298/rusentj.35.3.13
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Gorbunov OG, Efetov KA (2024) To the distribution of the genus Negotinthia O. Gorbunov, 2001 (Lepidoptera: Sesiidae) in the Crimea. Ecologica Montenegrina 73: 146–157. https://doi.org/10.37828/em.2024.73.15
Gorbunov OG, Efetov KA (2025a) To the distribution of Microsphecia brosiformis (Hübner, 1813) (Lepidoptera: Sesiidae) on the Crimean Peninsula (Russia). Ecologica Montenegrina 86: 1–14. https://doi.org/10.37828/em.2025.86.1
Gorbunov OG, Efetov KA (2025b) To the distribution of Conopia stomoxiformis (Hübner, 1790) (Lepidoptera: Sesiidae) on the Crimean Peninsula (Russia). Ecologica Montenegrina 87: 101–113. https://doi.org/10.37828/em.2025.87.5
Gorbunov OG, Efetov KA (2025c) To the distribution of Chamaesphecia (Scopulosphecia) schmidtiiformis (Freyer, 1836) (Lepidoptera: Sesiidae) on the Crimean Peninsula (Russia). Ecologica Montenegrina 87: 230–242. https://doi.org/10.37828/em.2025.87.11
Gorbunov OG, Efetov KA (2025d) To the distribution of Bembecia (Bembecia) ichneumoniformis ([Denis et Schiffermüller], 1775) (Lepidoptera: Sesiidae) on the Crimean Peninsula, Russia. Ecologica Montenegrina 90: 331–343. https://doi.org/10.37828/em.2025.90.14
Gorbunov OG, Efetov KA (2026a) To the morphology and distribution of Chamaesphecia (Chamaesphecia) astatiformis (Herrich-Schäffer, 1864) (Lepidoptera: Sesiidae) on the Crimean Peninsula. Ecologica Montenegrina 98: 46–61. https://doi.org/10.37828/em.2026.98.5
Gorbunov OG, Efetov KA 2026b. New data on the distribution and sexual activity of males of Stygioides colchica (Herrich-Schäffer, 1851) (Lepidoptera: Cossidae) on the Crimean Peninsula. Ecologica Montenegrina 101: 61–70. https://doi.org/10.37828/em.2026.101.6
Gorbunov OG, Efetov KA (2026c) On the distribution of the subgenus Ductispina Căpușe, 1973, of the genus Bembecia Hübner, 1819, on the Crimean Peninsula, Russia. Ecologica Montenegrina 102: 174–191. https://doi.org/10.37828/em.2026.102.13
Gorbunov OG, Efetov KA, Ivanov AV (2025a) First record of Bembecia (Opacosphecia) eversmanni O. Gorbunov, 2024 (Lepidoptera: Sesiidae) from the Crimean Peninsula, Russia. Ecologica Montenegrina 91: 47–58. https://doi.org/10.37828/em.2025.91.7
Gorbunov OG, Ivanov AV (2026) A new species of the genus Afromelittia O. Gorbunov et Arita, 1997 (Lepidoptera: Sesiidae) from Saudi Arabia. Ecologica Montenegrina 97: 99–109. https://doi.org/10.37828/em.2026.97.9
Gorbunov OG, Koshkin ES (2026) New data on clearwing moths (Lepidoptera: Sesiidae) of the tribe Synanthedonini in Khabarovsk Krai, with description of a new species. Russian Entomological Journal 35(2): 222–234. https://doi.org/10.15298/rusentj.35.2.13
Gorbunov OG, Snegovaya N, Shapoval NA, Mammadov IB-O, Yakovlev RV (2025b) Towards the knowledge of the fauna of clearwing moths (Lepidoptera, Sesiidae) of Nakhichevan Autonomous Republic, Azerbaijan. Acta Biologica Sibirica 11: 1379–1393. https://doi.org/10.5281/zenodo.17908580
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How to cite this article:
Gorbunov OG, Efetov KA (2026) To the morphology and distribution of Synanthedon andrenaeformis (Laspeyres, 1801) (Lepidoptera: Sesiidae) on the Crimean Peninsula, Russia. Acta Biologica Sibirica 12: 1471–1489. https://doi.org/10.5281/zenodo.24036288