Abstract
Here we present additional data and information about new records of Lepidoptera, Papilionoidea from the Republic of Khakassia and the southern Krasnoyarsk Kray, based on collections and observations in 1986−2025. Five species are reported for the fauna of these regions for the first time and fifteen are reliably confirmed. In addition, we verified the presence of a number of species that were not in the Catalogue for this region, but were noted by researchers in the past. A separate task of this work was an attempt to compile the most comprehensive bibliography of scientific works concerning the diurnal butterflies of Khakassia and the southern Krasnoyarsk Kray. A common eastward range shift pattern has been observed for several nemoral butterfly species, which we attribute to the onset of another Holocene Climatic Optimum.
References
Churkin SV, Bogdanov PV (2003) Two new subspecies of the genus Boloria Moore (Lepidoptera, Nymphalidae) from Siberia and Mongolia. Helios 4: 217–223.
Dragan SV (2003) Information on the collection of butterflies (Lepidoptera, Diurna) from the Republic of Khakassia, stored in the Zoological Museum of Khakassian State University named after N. F. Katanov. Collection of Khakass State University named after N.F. Katanov, Abakan. Ecology of Southern Siberia and Adjacent Territories 1: 73–75. [In Russian]
Dragan SV (2004) Lepidoptera, Diurna the mountain taiga belt of the central part of the Shaman Range (Western Sayan). Ecology of Southern Siberia and Adjacent Territories. Materials of the International Scientific School-Conference of Students and Young Scientists 1: 77. [In Russian]
Dragan SV (2005) On the occurrence of Carcharodus flocciferus (Zeller, 1847) (Lepidoptera, Hesperiidae) on the northern macroslope of the Western Sayan Mountains. Altai: Ecology and Nature Management. Proceedings of the IV Russian-Mongolian Scientific Conference of Young Scientists and Students. The Altai Region: Ecology and Nature Management. RIO BSPI named after V. M. Shukshin, Biysk, 119–120. [In Russian]
Dragan SV (2007) Lepidoptera Diurna as a Component of Biocenoses of the North-western Macroslope of Western Sayan. Ecology of Southern Siberia and Adjacent Territories 11(1): 93–95. [In Russian]
Dragan SV, Listvyagova NA (2016) On the Insect Fauna (Insecta) of the Khol-Bogaz Section of the Khakassky State Nature Reserve. Scientific Research in Nature Reserves and National Parks of Southern Siberia 6: 11–13. [In Russian]
Dragan SV (2011) New records of Leptidea reali Reissinger, 1989 (Lepidoptera, Pieridae) in the Southern Siberia. Euroasian Entomological Journal 10(1): 31–32. [In Russian]
Dragan SV (2018) First record of Apatura iris (Linnaeus, 1758) (Lepidoptera: Nymphalidae) from the Republic of Khakassia (South Siberia). Acta Biologica Sibirica 4(4): 114–115. https://doi.org/10.14258/abs.v4.i4.4884
Dragan SV, Isaeva IL, Sankina NI, Shuryshev EY (2019) Insects of the Oglakhty section of the Khakassky Nature Reserve. In: Natural Complex and Biodiversity of the Oglakhty Section of the Khakassky Nature Reserve. Khakassian Book Publishing House, Abakan, 197–205. [In Russian]
Dragan SV, Bezrukikh AV (2021) Contribution to the knowledge of the entomo-fauna of the Natural Reserve Iyusskij and its surroundings. In: Human-Nature Interaction in Specially Protected Natural Areas. Novokuznetsk, 43–47. [In Russian]
Dubatolov VV, Kosterin OE (2000) Nemoral species of Lepidoptera (Insecta) in Siberia: a novel view on their history and the timing of their range disjunctions. Entomologica Fennica 11: 141–166. https://doi.org/10.33338/ef.84061
Ermolajev WN (1935) Lepidopteren – Sammlungen aus verschiedenen Gegenden von Zentralsibirien. Folia Zoologica et hydrobiologica 7(2): 152–160, Abb, 1–3. Gorbunov PY, Kosterin OE (2003) The Butterflies (Hesperioidea and Papilionoidea) of North Asia (Asian part of Russia) in Nature. Vol. I. Rodina and Fodio, Gallery Fund, Moscow, Cheliabinsk, 392 pp.
Gorbunov PY, Kosterin OE (2007) The Butterflies (Hesperioidea and Papilionoidea) of North Asia (Asian part of Russia) in Nature. Vol. II. Rodina and Fodio, Aidis Producer's House, Moscow, 408 pp.
Haanshus K (1920) Lepidoptera ab Anders Orvin lecta. Macrolepidoptera. In: Insecta, ex-Sibiriameredionali et Mongolia, in itinere Orjan Olsen 1914 collecta. Norsk Entomologisk Tidsskrift 1: 10–17.
Jakobson AG (1901) Travel to Abakan Mountains. Trudy Russkogo entomologicheskogo obshestva 35(1–2): 5–14. [In Russian]
Ivonin VV, Nikolaev SL (2008a) Updated checklist of butterfly (Lepidoptera, Diurna) species of Tyva Republic. Altai Zoological Journal 2: 102–116. [In Russian]
Ivonin VV, Nikolaev SL (2008b) Data on geographical distribution of butterflies (Lepidoptera, Diurna) of Tyva Republic as revealed by expeditions of 2001–2005. Altai Zoological Journal 2: 117–121. [In Russian]
Ivonin VV, Kosterin OE, Nikolaev SL (2016) Butterflies (Lepidoptera, Diurna) of Novosibirskaya Oblast’, Russia. Nymphalidae, Satirinae. Euroasian Entomological Journal 15(2): 143–158. [In Russian]
Ivonin VV, Kosterin OE, Nikolaev SL, Yudina MA (2018) Butterflies (Lepidop-tera, Diurna) of Novosibirskaya Oblast’, Russia. 5. Updates and general discussion. Euroasian Entomological Journal 17(1): 25–52. [In Russian] http://doi.org/10.15298/euroasentj.17.1.04
Kondakov YP, Baranchikov YN (1975) Fauna and Ecological Principles of Conservation of Lepidoptera, Rhopalocera in Central Krasnoyarsk Territory. In: Protection and Rational Use of Forests in the Krasnoyarsk Territory. Institute of Forest and Wood, Siberian Branch of the USSR Academy of Sciences, Krasnoyarsk, 159–178. [In Russian]
Korb SK (2005) A Catalogue of the ex-USSR with remarks on systematics and nomenclature. Nizhny Novgorod, 158 pp.
Korb SK, Bolshakov LV (2011) Taxonomic notes upon the genus Euphydryas Scud-der, 1872 (Lepidoptera: Nymphalidae). Eversmannia 25–26(6): 25–33. [In Russian]
Korshunov YP (1969) The Butterflies of (Lepidoptera, Diurna) “Stolby” Nature Reserve in Mana District. Works of the State Nature Reserve "Stolby" [Trudy gosudarstvennogo zapovednika “Stolby”] 7: 165–203. [In Russian]
Korshunov YP (1972) Catalogue of diurnal butterflies (Lepidoptera, Rhopalocera) of the fauna of the USSR. Entomological Review [Entomologicheskoe Obozrenie] 51(1): 136–154. [In Russian]
Korshunov YP (1972) Catalogue of diurnal butterflies (Lepidoptera, Rhopalocera) of the fauna of the USSR. II. Entomological Review [Entomologicheskoe Obozrenie] 51(2): 352–368. [In Russian]
Korshunov YP (1978) Ecological and faunistic groups of butterflies (Lepidoptera, Diurna) in mountains of South Siberia. In: Arthropods of Siberia. Vol. 34. Nau-ka, Novosibirsk, 168–183. [In Russian]
Korshunov, YP (1984) New subspecies of unpaired fritillary Argynmis sagana Dbl, from Siberia. Insects and a helmints, new and little knoun species of the fauna of Siberia 17: 58–62. [In Russian]
Korshunov YP (1988) New butterflies (Lepidoptera, Rhopalocera) from Khakasia, Tuva and Yakutia. In: The taxonomy of animals of the Siberia. New and little known species of the fauma of Siberia 20: 65–80. [In Russian]
Korshunov YP, Gorbunov PY (1995) The Butterflies of Asian part of Russia. Ural University Press, Ekaterinburg, 202 pp. [In Russian]
Korshunov YP (2000) Butterflies of Ural, Siberia and Far Fast (keys and details). ООО "Vitel", Novosibirsk, 219 pp. [In Russian]
Korshunov YP, Nikolaev SL (2002) Toward the knowledge of the genus Oeneis Hübner, 1819 (Lepidoptera, Satyridae) of North Asia. Euroasian Entomological Journal 1(2): 147–172. [In Russian]
Koshantshikov WD (1923) Materialjen zur Macrolepidopteren fauna des Mi-nussinsk Bezirks. (Sibirien, Ienissei Gov.) Jarbuch des Martjanovischen Staats-museums in Minussinsk (Siberien), I, (I), I–VII & 1–50. [In Russian and German]
Koshantshikov WD (1924) Materialjen zur Macrolepidopteren fauna des Mi-nussinsk Bezirks. (Sibirien, Ienissei Gov.) Jarbuch des Martjanovischen Staats-museums in Minussinsk (Siberien), II, (I): 66–75. [In Russian and German]
Koshantshikov WD (1926) Materialjen zur Macrolepidopteren fauna des Mi-nussinsk Bezirks IV. Jarbuch des Martjanovischen Staatsmuseums in Minussinsk (Siberien), IV, (I): 79–82. [In Russian and German]
Koshantshikov WD (1928) Materialjen zur Macrolepidopteren fauna des Mi-nussinsk Bezirks VI. Jarbuch des Martjanovischen Staatsmuseums in Minussinsk (Siberien), VI, (I): 64–82. [In Russian and Germany]
Kosterin OE (2000) Description of a new subspecies of the Boloria pales ([Denis et Shiffermuller],1775) sensu Warren (1944) (Lepidoptera, Nymphalidae) from Altai. Far Eastern Entomologist 86: 1–10.
Kosterin OE (2023) Early summer aspect of butterflies (Lepidoptera: Papilionoidea) of Republic of Khakassia as examined in 2000, with some additional data. Amurian Zoological Journal 15(2): 325–354. https://www.doi.org/10.33910/2686-9519-2023-15-2-325-354
Larionov AV (2006) Butterflies in vacinity Ustinkino village in Ordzhonikidze District of Khakassia Republic. Reports on the 10th International Conference of Students and Young Scientists: Ecology of Southern Siberia and adjacent territories, 89 p. [In Russian]
Loshchev SM (2015) Entomofauna of the "Stolby" Nature Reserve. Proceedings of the "Stolby" State Nature Reserve [Trudy gosudarstvennogo zapovednika "Stolby"]. Issue 21. Krasnoyarsk, 216 pp. [In Russian]
Loshchev SM, Maksimov RE (2021) New Lepidoptera species (Insecta, Lepidoptera) in the Krasnoyarsk Pillars National Park, the south of the Krasnoyarskiy Kray and Khakassia. Scientific research in nature reserves and national parks of southern Siberia 10: 23–27. [In Russian]
Lukhtanov V, Lukhtanov A (1994) Die Tagfalter Nordwestasiens (Butterflies of Nord-West Asia). Herbipoliana. Buchreihe zur Lepidopterologie. Band 3. Dr. Eitschberger U (Ed.). Marktleuthen, 440 pp.
Lukhtanov VA (1989) A Review of Palaearctic Satyrids of the Genus Oeneis Hubner, 1816 (Lepidoptera, Saryridae). I. Vestnik Zoologii 2: 29–34. [In Russian]
Lukhtanov VA, Eitschberger UFJ (2000) Schmetterlinge der erde, Tagfalter. Teil 11. Nymphalide V. Oeneis. E. Bauer, T. Frankenbach. T Schmetterlinge der Erde. Goecke and Evers, Kaltern, 1–12, taf. A–B, 1–26. [In German]
Lukhtanov VA, Eitschberger UFJ (2001) Catalogue of the genera Oeneis and Davidiana. In: Bauer E, Frankenbach T (Eds) Butterflies of the world. Supplement 4. Goecke and Evers, Kaltern, 1–37.
Maksimov RE, Ivanov MA, Knyazev SA (2019) New finds of Heterocera (Insecta, Lepidoptera) in the Republic of Khakassia and on the South of Krasnoyarsk Region (South Siberia, Russia). Acta Biologica Sibirica 5(4): 12–20. http://doi.org/10.14258/abs.v5.i4.6803
Maksimov RE, Knyazev SA, Matov AYu, Makhov IA, Lostchev SM, Ivanov MA (2022) Additions to the fauna of Heterocera (Insecta, Lepidoptera) of the Republic of Khakassia and of the south of Krasnoyarsk Region (South Siberia, Russia) with a comparison of the moth flight timing after 100 years of W. Kozhantshikov's research. Acta Biologica Sibirica 8: 507–520. https://doi.org/10.5281/zenodo.7710474
Maksimov RE (2022) Wasily Dmitrievich Kozhanchikov. The Minusinsk Years (1915–1929). Scientific Notes of N.M. Martyanov Museum 4: 107–133. [In Russian]
Maksimov RE (2024) Wasily Dmitrievich Kozhanchikov. Additions and Corrections. Scientific Notes of N.M. Martyanov Museum 6: 122–148. [In Russian]
Maksimov RE (2025) The Norwegian Footprint in Southern Siberian Entomology. Tribune of Katanov Khakassian State University 4: 54, 27–34. [In Russian, with English summary]
Meinhard AA (1912) Review of Entomological Collections made in the Western Sayan Mountains and on the Enisei river from August 5 to September 8, 1903. Typo-Lithography of the Siberian Printing Association, Tomsk, 15 pp. [In Russian]
Pavel J (1901) Lepidopteren. In: Dritte Asiatische Forschungsreise des Grafen Eu-gen Zichy 2: 171–177.
Staudinger O (1886) Centralasiatische Lepidopteren. Entomologische Zeitung (Stettin) 47: 193-215, 225–256.
Staudinger O (1887) Centralasiatische Lepidopteren. Entomologische Zeitung (Stettin) 48: 49–102.
Staudinger O (1899) Ueber die Arten und Formen d en der Lycaena damon-Gruppe, Deutsche entomologische Zeitschrift Iris 12 (1): 137–155.
Staudinger O, Rebel H (1901) Macrolepidoptera. Catalog der Lepidopteren des palaearctischen Faunengebietes. Friedlander & Sohn, Berlin, 1: xxxii + 411 pp. https://doi.org/10.5962/bhl.title.120482
Sheljuzhko LA (1928a) Neue palearktische Lycaeniden. Lepidopterologische Rundschau 2(4): 44–46, 2(5): 51–52, 2(6): 62–64.
Sheljuzhko, LA (1928b) Bemerkungen über einige Lycaeniden des Bezirkes von Minussinsk (Gouvern. Jenissej, Sibirien). Lepidopterologische Rundschau 2(10): 110–112, 2(11): 116–120, 2(12): 128–132.
Sinev SYu (Ed.) (2008) Catalogue of the Lepidoptera of Russia. KMK Scientific Press Ltd., St. Petersburg–Moscow, 424 pp. [In Russian]
Sinev SYu (Ed.) (2019) Catalogue of the Lepidoptera of Russia. Second edition. Zoo-logical Institute RAN, St. Petersburg, 448 pp. [In Russian]
Sushkin PP, Chetverikov SS (1907) Lepidoptera of Minusinsk District, West Sayan and western part of Uryanhayskaya steppe. Proceedings of the Russian Entomology Society XXXVIII: 3–34. [In Russian]
Trybom F (1877) Dagfjärilar insamlade af svenska expeditionen till Jenisei 1876. Öfversigt af Kongl. Vetenskaps Akademiens Förhandlinder 34 (6): 35–51.
Tshikolovets VV, Yakovlev VR, Kosterin OE (2009) Butterflies of Altai, Sayans and Tuva (South Siberia). Tshikolovets Press, Padrubice, 374 pp.
Tschugunov SM (1912) Cheshuekrylye, sobrannye letom 1909 na ozere Ingol, Enisseyskoy gubernii. Russkoye Entomologicheskoye Obozrenye XII(2): 216–219. [In Russian]
Tschugunov SM (1913) Cheshuekrylye, sobrannye letom 1912 v Minussinskom uezde Enisseyskoy gubernii. Russkoye Entomologicheskoye Obozrenye XIII(2): 360–367. [In Russian]
Tschugunov SM (1916) Contribution à la faune de Lepidoptères des prémonts de l'Alataou de Kuznetzk. Revue Russe d`Entomologie 16(1/2): 75–105.
Tuzov VK, Bogdanov PV, Devyatkin AL, Kaabak LV, Korolev VA, Murzin VS, Samodurov GD, Tarasov EA (1997) Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera), 1. Pensoft, Sofia-Moscow, 480 pp.
Tuzov VK, Bogdanov PV, Churkin SV, Dantchenko AV, Devyatkin AL, VA, Murzin VS, Samodurov GD, Zhdanko AB (2000) Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Rhopalocera), 2. Pensoft, Sofia-Moscow, 600 pp.
Tuzov VK (2015) Bibliography on the butterflies of Russia and adjacent countries (1758-2008). Tshikolovets Press, Padrubice, 682 pp.
Ustjuzhanin PYa, Maksimov RE (2023) The Pterophoridae fauna of Khakassia Republic and south of Krasnoyarsk region (Russia) (Insecta: Lepidoptera). SHILAP Revista de lepidopterologia. 51(204): 581−591. https://doi.org/10.57065/shilap.781
Ustjuzhanin, PYa, Kovtunovich VN, Maksimov RE (2024) New data on the plume moths from Khakassia and the South of the Krasnoyarsk Territory (Lepidoptera: Pterophoridae). Ecologica Montenegrina 80: 13−20. https://doi.org/10.37828/em.2024.80.2
Wnukovsky WW (1929–1930) Révision des quelques articles par A. Meinhard et par S. Tshougounow (S. Cugunov) à la faune des Lépidoptères des Altai, pays de Minousinsk, monts Sajan et de Mongolie septentrionale-occidentale. Trudy Sibirskogo instituta selskogo khozyajstva i lesovodstva 13(1/2): 275–285. [In Russian]
Yanovsky VM (1996) Forest entomofauna of the Sayano-Shushensky Biosphere Reserve. Institute of Forestry named after V.N. Sukachev, Siberian Branch of the Russian Academy of Sciences, 1–47. [In Russian]
Yanovsky VM (2002) To the fauna of forest insects of the Achinsk forest-steppe (Krasnoyarsk Kray). In: Baranchikov YuI, Petrenko ES (Eds) Entomological research in Siberia 2: 118–130. [In Russian]

This work is licensed under a Creative Commons Attribution 4.0 International License.
