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Submission Preparation Checklist

As part of the submission process, authors are required to check off their submission's compliance with all of the following items, and submissions may be returned to authors that do not adhere to these guidelines.
  • The text, if submitted to a peer-reviewed section (e.g., Articles), has had the authors' names removed. If an author is cited, "Author" and year are used in the bibliography and footnotes, instead of author's name, paper title, etc. The author's name has also been removed from the document's Properties, which in Microsoft Word is found in the File menu.
  • The submission file is in Microsoft Word or RTF document file format.
  • The text adheres to the stylistic and bibliographic requirements outlined in the Author Guidelines, which is found in Submissions.
  • The text is single-spaced; uses a 11-point font; employs italics, rather than underlining (except with URL addresses); and all illustrations, figures, and tables are placed within the text at the appropriate points, rather than at the end.
  • All URL addresses in the text (e.g., are activated and ready to click
  • The submission has not been previously published, nor is it before another journal for consideration (or an explanation has been provided in Comments to the Editor).

Author Guidelines

The journal publishes original, previously unpublished articles on the following fields:

  • Faunistics
  • Floristics
  • Biogeography
  • Biological systematics
  • Nature conservation and protected areas
  • Reviews of published articles and monographs on the above topics
  • Memorial articles

We accepted the manuscripts exclusively in English since 2020. There are no restrictions on the article volume and number of articles in the issue. The journal is published four times a year in the “continuous publication” mode.

You can uploaded your ms through OJS interface or send it directly to Editorial on

In the fields of faunistics and floristics, we accept articles of two types: floral and faunal lists on any region of the world (areas, regions, countries, mountain ranges, national parks), and faunal and floral discoveries (finding new species for the regions, additions to previously published inventories…). We also accept articles on methodology of faunal and floral studies. The materials up to three pages are placed in the category Short Communication.

The journal policy includes a mandatory procedure of blind peer review. Preference is given to articles illustrated with distribution maps, photographs of biotops, biological objects in nature and collection specimens.

  1. The following structure of articles is recommended: Introduction. Material and methods. Results. Discussion. Acknowledgments. References.
  2. The article title must be given in full and short form (full title and running head). All the authors must be listed with current contact information including the address, place of work and e-mail, the corresponding author is marked with (*). The abstract should not exceed 500 characters, the keywords are up to 10. For faunal and floral discoveries the given structure (Introduction, Materials and Methods, Results, Conclusion, Acknowledgements, References) can be changed, reflecting only factology. References should be made in accordance with ZooKeys journal.
  3. Texts in English should be verified by a native speaker. Latin words for species and genera must be italicized. At the first mention of a taxon its full title including author and year of description should be given (if necessary – in brackets): Papilio machaon Linnaeus, 1758. The subsequent references to the taxon should be given in a shortened version – P. machaon. In the description of new taxa, authors should strictly follow the Codes of botanical and zoological nomenclature.
  4. The literature should be cited as recommended below and also please refer to The references must be arranged alphabetically by the last name of the first author or, if author is not available - by title.
  5. It is advisable to adhere to the following design of material for faunal and floral lists (discoveries). Be sure to mention the collection or herbarium, where the material is deposited. The legend to abbreviations may be given in the Materials and Methods section.

Material examined. 3 males, SW Mongolia, Hovd Aimak, Dzhungarian Gobi desert, 20 km N Zeegijn border post  (45°17’N / 92°09’E), h= 1400 м, 29-30.06.2015, leg. R. Yakovlev (coll. R. Yakovlev, Barnaul).

Material examined. 1 female, Kwango, Kikwit [5°02’00” S / 18°49’00”E], 1938, Rev. F. Placide (MRAC). At the mention of the geographical origin labels without coordinates it is desirable to give the coordinates of the area in brackets.

Material examined. «Тувинская АССР, Монгун-Тайгинский р-н, верх. р. Мугур, 17-19. VIII. 1978. А.С. Ревушкин, В.В. Хлопов, Т.В. Жигальцова» [Russia, Tuva Rep., Mongun-Taiga distr., upper stream of Mugur River, 17−19.08.1978, A.S. Revushkin, V.V. Khlopov, T.V. Zhigal’tsova]. The labels can be given in the original language in commas and translated into English in square brackets.

      6. Graphs and diagrams should be created in a specialized editor included into MS Word or in vector graphics editor: Corel Draw, Adobe Illustrator (.cdr, .ai, .wmf, the text inside graph should not be “saved as vector”). Use of non-standard fonts is undesirable. Raster versions, as well as graphs and charts made "by hand', are not permitted. Graphs and diagrams created in MS Excel are undesirable. Diagrams should be in black and white, all the divisions - marked with large hatching legible when printed.

       7. Illustrations should be presented in LineArt (raster, black and white) — TIFF 600–1200 dpi (depending on details), LZW compression. Grey (photo) — JPEG 300–600 dpi (compression ratio 8–10). Vector drawings should be given in EPS, AI, CDR, not using non-standard fills and fonts. All raster images must have real, not interpolated resolution. Images created in non-graphic editors, as well as illustrations imported into Word, are not permitted.

       8. Titles of tables and figure captions are given in the text of the article. Files with illustrations should be named as follows: petrov_1, petrov_2, petrov_3_12… Figure captions should be presented as follows:

Fig. 1. Paralophonotus auroguttata (Herrich-Schaff er, [1854]), male, Congo, Baraka in Kivu Province, Lt. Ghesquiere (MRAC).

Fig. 1. Map of collecting localities: 1. Hovd Aimak, Dzun-Dzhargalant-Khairkhan, Ar-Shatyn-Gol river Valley (47°44’N; 92°27’E); 2. Hovd Aimak, Dzhungarian Gobi desert, Uvkhod-Ula (Ovkhood-Uul) Mt. (45°48’N; 91°06’E); 3. Hovd Aimak, Dzhungarian Gobi desert, 20 km N Zeegijn border post  (45°17’N; 92°09’E); 4. Hovd Aimak, Bajtag-Bogd-Uul Mts., Baruun-Khargaityn-Gol river Valley (45°16’N; 90°57’E); 5. Hovd Aimak,  Ikh-Khavtag-Nuruu Mts., near Zhugentijn spring, (45°04’N; 92°12’E); 6. Hovd Aimak, Ikh-Khavtag-Uul Mts., near Yargajtyn-Sajr gorge (45°03’N; 92°09’E).

Fig. 7. Map of distribution of Hyles svetlana Shovkoon, 2010 and north-western Chinese records of Eremurus inderiensis (Steven) Regel (from Pittaway & Kitching, 2016 with additions).

Figures 1−7. Aholcocerus, adults and male genitalia: 1. Aholcocerus jakli Yakovlev & Witt sp. nov., holotype (MWM); 2. Aholcocerus ihleorum Yakovlev & Witt, 2009, holotype (MWM); 3. Aholcocerus ronkayorum Yakovlev, 2006, holotype (MWM); 4. Aholcocerus sevastopuloi Yakovlev, 2011, holotype (BMNH); 5. Aholcocerus verbeeki (Roepke, 1957), holotype (RMNH); 6. Aholcocerus verbeeki (Roepke, 1957), female, allotype (RMNH); 7. Aholcocerus jakli Yakovlev & Witt sp. nov., male genitalia

        9. Tables should be submitted apart from the article text on separate pages after the list of references. Avoid multi-page tables; it is preferable to divide large data into several tables. You can mark the place in the text where the table is to be inserted [Table x]. Tables should be numbered in Arabic numerals and have headlines in Russian and English, briefly revealing their content. Subtitles of columns should be concise and informative. Units of measurement should be indicated after the comma. In tables, don't use vertical and horizontal lines separating columns. Additional information to clarify the content of the table should be placed under it as a note.

10. Cited literature should be restricted to published papers. Only papers that have been accepted for publication may be incorporated, with the notation 'in press' (and the journal in which it will appear). Unpublished data, manuscripts in preparation and unpublished papers should be noted as 'pers. comm.' or 'unpubl. data'. Check every reference in the text of the article with the list of used literature. All the names of periodicals should be checked thoroughly. In-text references consist of the surname of the author or authors and the year of publication of the document. End references are unnumbered and appear in alphabetical order by author and year of publication, with multiple works by the same author listed in chronological order.

Each example of an end reference is accompanied here by an example of a corresponding in-text reference. For more details and many more examples, see Chapter 29 of Scientific Style and Format.


For the end reference, list authors in the order in which they appear in the original text. The year of publication follows the author list. Use periods to separate each element, including author(s), date of publication, article and journal title, and volume or issue information. Location (usually the page range for the article) is preceded by a colon.

Author(s). Date. Article title. Journal title. Volume(issue):location.

Journal titles are generally abbreviated according to the List of Title Word Abbreviations maintained by the ISSN International Centre. See Appendix 29.1 in Scientific Style and Format for more information.

For the in-text reference, use parentheses and list author(s) by surname followed by year of publication.

(Author(s) Year)

For articles with 2 authors, names are separated by a comma in the end reference but by “and” in the in-text reference.

Mazan MR, Hoffman AM. 2001. Effects of aerosolized albuterol on physiologic responses to exercise in standardbreds. Am J Vet Res. 62(11):1812–1817.

(Mazan and Hoffman 2001)

For articles with 3 to 10 authors, list all authors in the end reference; in the in-text reference, list only the first, followed by “et al.”

Smart N, Fang ZY, Marwick TH. 2003. A practical guide to exercise training for heart failure patients. J Card Fail. 9(1):49–58.

(Smart et al. 2003)

For articles with more than 10 authors, list the first 10 in the end reference, followed by “et al.”

Pizzi C, Caraglia M, Cianciulli M, Fabbrocini A, Libroia A, Matano E, Contegiacomo A, Del Prete S, Abbruzzese A, Martignetti A, et al. 2002. Low-dose recombinant IL-2 induces psychological changes: monitoring by Minnesota Multiphasic Personality Inventory (MMPI). Anticancer Res. 22(2A):727–732.

(Pizzi et al. 2002)

Volume with no issue or other subdivision

Laskowski DA. 2002. Physical and chemical properties of pyrethroids. Rev Environ Contam Toxicol. 174:49–170.

(Laskowski 2002)

Volume with issue and supplement

Gardos G, Cole JO, Haskell D, Marby D, Paine SS, Moore P. 1988. The natural history of tardive dyskinesia. J Clin Pharmacol. 8(4 Suppl):31S–37S.

(Gardos et al. 1988)

Volume with supplement but no issue

Heemskerk J, Tobin AJ, Ravina B. 2002. From chemical to drug: neurodegeneration drug screening and the ethics of clinical trials. Nat Neurosci. 5 Suppl:1027–1029.

(Heemskerk et al. 2002)

Multiple issue numbers

Ramstrom O, Bunyapaiboonsri T, Lohmann S, Lehn JM. 2002. Chemical biology of dynamic combinatorial libraries. Biochim Biophys Acta. 1572(2–3):178–186.

(Ramstrom et al. 2002)

Issue with no volume

Sabatier R. 1995. Reorienting health and social services. AIDS STD Health Promot Exch. (4):1–3.

(Sabatier 1995)


In the end reference, separate information about author(s), date, title, edition, and publication by periods. The basic format is as follows:

Author(s). Date. Title. Edition. Place of publication: publisher. Extent. Notes.

Extent can include information about pagination or number of volumes and is considered optional. Notes can include information of interest to the reader, such as language of publication other than English; such notes are optional. Essential notes provide information about location, such as a URL for online works. See Chapter 29 for more information.

For books with 2 authors, names are separated by a comma in the end reference but by “and” in the in-text reference.

Leboffe MJ, Pierce BE. 2010. Microbiology: laboratory theory and application. Englewood (CO): Morton Publishing Company.

(Leboffe and Pierce 2010)

For books with 3 to 10 authors, list all authors in the end reference; in the in-text reference, list only the first, followed by “et al.”

Ferrozzi F, Garlaschi G, Bova D. 2000. CT of metastases. New York (NY): Springer.

(Ferrozzi et al. 2000)

For books with more than 10 authors, list the first 10 in the end reference, followed by “et al.”

Wenger NK, Sivarajan Froelicher E, Smith LK, Ades PA, Berra K, Blumenthal JA, Certo CME, Dattilo AM, Davis D, DeBusk RF, et al. 1995. Cardiac rehabilitation. Rockville (MD): Agency for Health Care Policy and Research (US).

(Wenger et al. 1995)

Organization as author

[ALSG] Advanced Life Support Group. 2001. Acute medical emergencies: the practical approach. London (England): BMJ Books.

(ALSG 2001)

Author(s) plus editor(s) or translator(s)

Klarsfeld A, Revah F. 2003. The biology of death: origins of mortality. Brady L, translator. Ithaca (NY): Cornell University Press.

Luzikov VN. 1985. Mitochondrial biogenesis and breakdown. Galkin AV, translator; Roodyn DB, editor. New York (NY): Consultants Bureau.

(Klarsfeld and Revah 2003)

(Luzikov 1985)

Chapter or other part of a book, same author(s)

Gawande A. 2010. The checklist manifesto: how to get things right. New York (NY): Metropolitan Books. Chapter 3, The end of the master builder; p. 48–71.

(Gawande 2010)

Chapter or other part of a book, different authors

Rapley R. 2010. Recombinant DNA and genetic analysis. In: Wilson K, Walker J, editors. Principles and techniques of biochemistry and molecular biology. 7th ed. New York (NY): Cambridge University Press. p. 195–262.

(Rapley 2010)

Multivolume work as a whole

Alkire LG, editor. 2006. Periodical title abbreviations. 16th ed. Detroit (MI): Thompson Gale. 2 vol. Vol. 1, By abbreviation; vol. 2, By title.

(Alkire 2006)

Dissertations and Theses

Lutz M. 1989. 1903: American nervousness and the economy of cultural change [dissertation]. [Stanford (CA)]: Stanford University.

(Lutz 1989)


Blanco EE, Meade JC, Richards WD, inventors; Ophthalmic Ventures, assignee. 1990 Nov 13. Surgical stapling system. United States patent US 4,969,591.

(Blanco et al. 1990)


Weiss R. 2003 Apr 11. Study shows problems in cloning people: researchers find replicating primates will be harder than other mammals. Washington Post (Home Ed.). Sect. A:12 (col. 1).

(Weiss 2003)


Indicate a copyright date with a lowercase “c”.

Johnson D, editor. c2002. Surgical techniques in orthopaedics: anterior cruciate ligament reconstruction [DVD]. Rosemont (IL): American Academy of Orthopaedic Surgeons. 1 DVD.

(Johnson c2002)

Websites and Other Online Formats

References to websites and other online formats follow the same general principles as for printed references, with the addition of a date of update/revision (if available) along with an access date and a URL.


Format for end reference:

Title of Homepage. Date of publication. Edition. Place of publication: publisher; [date updated; date accessed]. Notes.

If no date of publication can be determined, use a copyright date (if available), preceded by “c”. Include the URL in the notes.

APSnet: plant pathology online. c1994–2005. St Paul (MN): American Phytopathological Association; [accessed 2005 Jun 20].

For the in-text reference, include only the first word or two of the title (enough to distinguish it from other titles in the reference list), followed by an ellipsis.

(APSnet . . . c1994–2005)

Online journal article

Format for end reference:

Author(s) of article. Date of publication. Title of article. Title of journal (edition). [date updated; date accessed];Volume(issue):location. Notes.

A DOI (Digital Object Identifier) may be included in the notes in addition to a URL, if available:

Savage E, Ramsay M, White J, Beard S, Lawson H, Hunjan R, Brown D. 2005. Mumps outbreaks across England and Wales in 2004: observational study. BMJ. [accessed 2005 May 31];330(7500):1119–1120. doi:10.1136/bmj.330.7500.1119.

(Savage et al. 2005)


Format for end reference:

Author(s). Date of publication. Title of book. Edition. Place of publication: publisher; [date updated; date accessed]. Notes.


Brogden KA, Guthmille JM, editors. 2002. Polymicrobial diseases. Washington (DC): ASM Press; [accessed February 28, 2014].

(Brogden and Guthmille 2002)


Format for end reference:

Author’s name. Date of publication. Title of post [descriptive word]. Title of blog. [accessed date]. URL.


Fogarty M. 2012 Aug 14. Formatting titles on Twitter and Facebook [blog]. Grammar Girl: Quick and Dirty Tips for Better Writing. [accessed 2012 Oct 19].

(Fogarty 2012)

Forthcoming or Unpublished Material

Not all forthcoming or unpublished sources are suitable for inclusion in reference lists. Check with your publisher if in doubt.

Forthcoming journal article or book

Journal article:

Farley T, Galves A, Dickinson LM, Perez MJ. Forthcoming 2005 Jul. Stress, coping, and health: a comparison of Mexican immigrants, Mexican-Americans, and non-Hispanic whites. J Immigr Health.

(Farley et al. 2005)


Goldstein DS. Forthcoming 2006. Adrenaline and the inner world: an introduction to scientific integrative medicine. Baltimore (MD): Johns Hopkins University Press.

(Goldstein 2006)

Paper or poster presented at meeting

Unpublished presentations are cited as follows:

Antani S, Long LR, Thoma GR, Lee DJ. 2003. Anatomical shape representation in spine x-ray images. Paper presented at: VIIP 2003. Proceedings of the 3rd IASTED International Conference on Visualization, Imaging and Image Processing; Benalmadena, Spain.

Charles L, Gordner R. 2005. Analysis of MedlinePlus en Español customer service requests. Poster session presented at: Futuro magnifico! Celebrating our diversity. MLA ’05: Medical Library Association Annual Meeting; San Antonio, TX.

(Atani et al. 2003)

(Charles and Gordner 2005)

References to published presentations are cited much like contributions to books, with the addition of information about the date and place of the conference. See Chapter 29 for more information.

Personal communication

References to personal communication are placed in running text rather than as formal end references. Permission is usually required and should be acknowledged in an “Acknowledgment” or “Notes” section at the end of the document.

. . . and most of these meningiomas proved to be inoperable (2003 letter from RS Grant to me; unreferenced, see “Notes”) while a few were not.


Please also refer to:

The author(s) can also use the article template for manuscript submission (download template)

Accepted manuscripts - The text of accepted manuscripts should be provided by e-mail, preferably as a Word for Windows document. Manuscripts are edited to enhance communication between author and reader. Proofs are provided as a PDF. In the proof stage only essential corrections can be made, and any additional alterations may be charged to the author. Corrected proofs should be returned within 1 week. Corresponding authors receive a complimentary copy of the issue in which the publication appears and an electronic version of the final publication (PDF).

Privacy Statement

Editors will treat all manuscripts submitted to Acta Biologica Sibirica in confidence. Reviewers are also required to treat manuscripts confidentially. Acta Biologica Sibirica will not share manuscripts with third parties outside of ABS except in cases of suspected misconduct.