A checklist of bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) from Siberia and the Russian Far East
Статьи
DOI: 10.5281/zenodo.7726527

A checklist of bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) from Siberia and the Russian Far East

Institute of Systematics and Ecology of Animals, SB RAS, Frunze street 11, Novosibirsk, 630091, Russia; Altai State University, Lenina 61, Barnaul, 656049, Russia

Аннотация

Currently, 185 species of the family Scolytidae and three species of the family Platypodidae are recorded from Asian Russia. In total, 99 species of bark beetles are found in Siberia and 168 species in the Russian Far East. Platypodidae are known anly from the south of the the Russian Far East. Two species of Scolytidae are found in Yamalo-Nenets Autonomous Okrug, 13 species in Khanty-Mansi Autonomous Okrug, 19 species in Tyumen Oblast, four species in Kurgan Oblast, three species in Omsk Oblast, 37 species in Tomsk Oblast, 28 species in Novosibirsk Oblast, 32 species in Kemerovo Oblast, 25 species in Altay Krai, 53 species in Altai Republic, 62 species in Krasnoyarsk Krai, 11 species in Republic of Khakassia, 24 species in Tyva Republic, 55 species in Irkutsk Oblast, 61 species in Buryatiya Republic, 40 species in Zabaikalskii Krai, 52 species in Sakha (Yakutia) Republic, 25 species in Kamchatka Oblast, one species in Chukotka Autonomous Okrug, 19 species in Magadan Oblast, 45 species in Amur Oblast, one species in Jewish Autonomous Oblast, 67 species in Khabarovsk Krai, 130 species in Primorsky Krai, 87 species in Sakhalin Is. and 68 species in Kuriles Isl.

Corresponding author: Andrei A. Legalov (fossilweevils@gmail.com)

Academic editor: R. Yakovlev | Received 2 November 2021 | Accepted 25 November 2022 | Published 14 December 2022

http://zoobank.org/33C16C41-D1F3-4B67-A3EC-F78E54C80460

Citation: Legalov AA (2022) A checklist of bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) from Siberia and the Russian Far East. Acta Biologica Sibirica 8: 623–646. https://doi.org/10.14258/abs.v8.e39

Keywords

Curculionoidea, beetles, fauna, distribution, North Asia, Russia.

Introduction

The first summarized data about the bark beetles was given by Heyden (1880-1881). The analytical reviews of scolytids were presented by Krivolutskaya (1983) and Yanovskij (1999). The keys to the bark and ambrosia beetles from the Russian Far East were compiled by Krivolutskaya (1996). In 2020, the author (Legalov 2020) published a list of Curculionoidea except the families Scolytidae and Platypodidae from Asian Russia. This is the first review of the bark and ambrosia beetles from Siberia and the Russian Far East.

Material and methods

The studied materials are deposited in the Institute of Systematics and Ecology of Animals (Novosibirsk), Zoological Institute (St. Petersburg), and some private collections.

The published records of the bark and ambrosia beetles from Siberia and Russian Far East (Heyden 1880–1881; Lavrov 1927; Kiseleva 1928, 1946, 1951, 1952; Kurentsov 1941; Eggers 1942; Florov 1949; Cherepanov 1952; Stark 1952; Krivolutskaya 1958, 1961, 1965a, 1965b, 1973, 1983, 1996a, 1996b; Rudnev 1958; Sokanovsky 1960; Lindeman 1961; Lurie and Lindeman 1961; Isaev and Tarasova 1965; Nemkova 1965; Galkin 1966; Ivliev and Kononov 1966, 1974; Opanassenko and Kononenko 1966; Krivolutskaya and Kupyanskaya 1970; Averensky 1971, 1999; Kononenko and Opanassenko 1971; Berezhnykh 1979; Nobuchi 1979; Gurov and Dryannykh 1982; Shatilov 1985, 1987; Wood and Bright 1992; Yanovskij 1999; Legalov and Sitnikov 2000; Mandelshtam 2001, 2002; Petrov and Mandelshtam 2002; Izhevsky et al. 2005; Krivets and Chemodanov 2005; Mandelshtam et al. 2007, 2018; Park and Lyu 2007; Bukhkalo et a. 2011, 2014; Kerchev 2011; Knížek 2011; Krivets et al. 2011; Krivets and Vysotina 2011; Petrov 2011, 2018; Akulov and Mandelshtam 2012a, 2012b; Efimov and Legalov 2012; Mandelshtam and Petrov 2016, 2019, 2022; Alonso-Zarazaga et al. 2017; Kerchev et al. 2019; Petrov et al. 2019; Johnson et al. 2020; Mandelshtam and Sergeev 2020; Petrov and Shamaev 2020; Agafonova et al. 2021; Shamaev 2021; Mandelshtam et al. 2022, etc.) are included.

The high systematics of studied taxa are from the works (Wood 1986, 1993; Zherichin and Egorov 1991; Morimoto and Kojima 2004; Gratshev and Legalov 2014; Legalov 2015).

Abbreviations for the names of federal subjects are follow (fig. 1): West Siberia: Yamalo-Nenets Autonomous Okrug - YAN, Khanty-Mansi Autonomous Okrug - KHM, Tyumen Oblast - TMN, Kurgan Oblast - KURG, Omsk Oblast - OMS, Tomsk Oblast - TOM, Novosibirsk Oblast - NOV, Kemerovo Oblast - KEM, Altay Krai - ALT, Altai Republic - RAL, Krasnoyarsk Krai - KRN, Republic of Khakassia - KHA, Tyva Republic - TUV, East Siberia: Irkutsk Oblast - IRK, Buryatiya Republic - BUR, Zabaikalskii Krai (formerly Chita Oblast) - CHT, Sakha (Yakutia) Republic - YAK, Far East: Kamchatka Oblast - KAM, Chukotka Autonomous Okrug - CHUK, Magadan Oblast - MAG, Amur Oblast - AMUR, EAO - Jewish Autonomous Oblast, Khabarovsk Krai - KHAB, Primorsky Krai - PRIM, Sakhalin Is. - SAKH, Kuriles Isl. - KUR.

Figure 1.Map of the administrative units of studied area.

Results

Superfamily Curculionoidea Latreille, 1802

Family Scolytidae Latreille, 1806

Subfamily Hylesininae Erichson, 1836

Tribe Hylastini LeConte, 1876

Genus Hylastes Erichson, 1836

ater (Paykull, 1800) – KHM, TMN, TOM, NOV, RAL, KRN, IRK, BUR, AMUR.

= angusticollis Eggers, 1929

brunneus Erichson, 1836 – TMN, TOM, NOV, KEM, ALT, RAL, KRN, BUR, CHT, YAK, KHAB, PRIM.

= aterrimus Eggers, 1933

cunicularius Erichson, 1836 (Fig. 2a) – KHM, TMN, TOM, RAL, KRN, IRK, BUR, YAK, PRIM, SAKH.

= starki Eggers, 1933

opacus Erichson, 1836 – TMN, TOM, NOV, ALT, RAL, KRN, IRK, BUR, CHT, YAK, AMUR, PRIM.

paralellus Chapuis 1875 – “East Siberia, Far East” [Knížek 2011].

obscurus Chapuis, 1875 – RAL, IRK, YAK, KAM, AMUR, KHAB, PRIM, SAKH.

= plumbeus Blandford, 1894

= septentrionalis Eggers, 1923

Genus Hylurgops Le Conte, 1876

glabratus (Zetterstedt, 1828) – KHM, TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

= paykulli Duftschmidt, 1825

= decumanus Erichson, 1836

= tenebrosus Sahlberg, 1836

interstitialis (Chapuis, 1875) – YAK, MAG, KAM, AMUR, KHAB, PRIM.

= imitator Reitter, 1900

longipilus Reitter, 1894 – IRK, BUR, KAM, AMUR, KHAB, PRIM, SAKH.

palliatus (Gyllenhal, 1813) – KHM, TMN, NOV, TOM, KEM, RAL, KHA, KRN, BUR, YAK, KHAB, PRIM, SAKH, KUR.

=parvus Eggers, 1933

spessivtsevi Eggers, 1914 – IRK, BUR, YAK, AMUR, KHAB, PRIM, SAKH.

transbaicalicus Eggers, 1941 – YAK, KAM, KHAB, PRIM.

Tribe Hylesinini Erichson, 1836

Genus Alniphagus Swaine, 1918

costatus (Blandford, 1894) (Fig. 2b) – KHAB, PRIM, SAKH, KUR.

= alni Niisima, 1909

Genus Hylesinus Fabricius, 1801

cholodkovskyi Berger, 1916 – PRIM.

cingulatus Blandford, 1894 – KHAB, PRIM.

eos Spessivtsev, 1919 – KHAB, PRIM.

laticollis Blandford, 1894 – KHAB, PRIM.

= striatus Eggers, 1933

= pravdini Stark, 1936

nobilis Blandford, 1894 – PRIM.

= shabliovskyi Kurenzov, 1941

mandshuricus Eggers, 1922 – PRIM.

tristis Blandford, 1894 (Fig. 2c)– PRIM.

= lubarskyi Stark, 1936

Genus Longulus Krivolutskaya, 1968

elatus (Niisima, 1913) (Fig. 2d) – KUR.

Tribe Hylurgini Gistel, 1848

Genus Dendroctonus Erichson, 1836

micans (Kugelann, 1794) (Fig. 2e) – KHM, KURG, NOV, ALT, RAL, KRN, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

Genus Hylurgus Latreille, 1806

ligniperda (Fabricius, 1787) – ALT.

Genus Tomicus Latreille, 1802

minor (Hartig, 1834) – TMN, NOV, TOM, RAL, KRN, IRK, BUR, YAK, AMUR, KHAB, PRIM.

pilifer (Spessivtsev, 1919) – KHAB, PRIM.

piniperda (Linnaeus, 1758) – TMN, NOV, TOM, KEM, ALT, RAL, KRN, IRK, BUR, YAK, AMUR, PRIM.

puellus (Reitter, 1894) – KHAB, PRIM, SAKH.

= puellus f. orientalis Krivolutskaya, 1956

= starki Eggers, 1929

Genus Xylechinus Chapuis, 1869

bergeri Spessivtsev, 1919 – PRIM

pilosus (Ratzeburg, 1837) – NOV, TOM, KEM, ALT, RAL, KHA, KRN, TUV, IRK, BUR, YAK, MAG, KAM, KHAB, PRIM, SAKH.

Tribe Hyorrhynchini Hopkins, 1915

Genus Pseudohyorrynchus Murayama, 1950

wadai Murayama, 1950 (Fig. 2f) – KUR.

= wadai kurilensis Krivolutskaya, 1956

Tribe Diamerini Hagedorn, 1909

Genus Sphaerotrypes Blandford, 1894

imitans Eggers, 1926 – PRIM.

juglansis Krivolutskaya, 1970 – PRIM.

Tribe Phloeotribini Chapuis, 1869

Genus Phloeotribus Latreille, 1797

spinulosus (Rey, 1883) – KEM, RAL, KRN, IRK, BUR, YAK, SAKH.

Tribe Polygraphini Chapuis, 1869

Genus Carphoborus Eichhoff, 1864

cholodkovskyi Spessivtsev, 1916 – NOV, TOM, ALT, RAL, KRN, IRK, BUR, CHT, YAK, MAG, AMUR.

jurinskii Eggers, 1910 – IRK, YAK.

rossicus Semenov, 1902 – KEM, KRN.

teplouchovi Spessivtsev, 1916 – NOV, KEM, ALT, RAL, KRN, IRK, BUR, CHT, YAK, MAG, AMUR.

Genus Polygraphus Erichson, 1836

abietis Kurentsov, 1941 – KHAB, PRIM.

gracilis Niisima, 1909 – KHAB, PRIM, SAKH, KUR.

horyurensis Murayama, 1937 – SAKH.

jezoensis Niisima, 1909 – KAM, KHAB, PRIM, SAKH.

kisoensis Niisima, 1941 – SAKH.

nigrielytris Niisima, 1913 – KAM, SAKH, KUR.

poligraphus (Linnaeus, 1758) – TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, KAM, AMUR, KHAB, PRIM, SAKH, KUR.

= pubescens Fabricius, 1792

= griseus Eggers, 1932

proximus Blandford, 1894 – NOV, TOM, KEM, ALT, RAL, KRN, KHA, IRK, KHAB, PRIM, SAKH, KUR.

= abietis Kurenzov, 1941

= horyurensis Murayama, 1937

= laticollis Eggers, 1926

punctifrons Thomson, 1886 – TOM, RAL, KRN, YAK, KAM, SAKH.

= seriatus Reitter, 1913

shariensis Niisima, 1941 – SAKH, KUR.

ssiori Niisima, 1909 – SAKH, KUR.

subopacus Thomson, 1871 – TOM, KEM, RAL, KRN, IRK, BUR, YAK, KAM, AMUR, KHAB, PRIM, SAKH, KUR.

= sachalinensis Eggers, 1926

Subfamily Scolytinae Latreille, 1806

Tribe Scolytini Latreille, 1806

Genus Scolytus Geoffroy, 1762

aratus Blandford, 1884 – AMUR, PRIM, SAKH, KUR.

= aequipunctatus Niisima, 1905

= brevipennis Kurenzov, 1941

= intermedius Kurenzov, 1941

butovitschi Stark, 1936 – KRN, BUR, PRIM.

= butovitschi Eggers, 1942

chikisanii Niisima, 1905 – PRIM, SAKH, KUR.

= curviventralis Niisima, 1905

= mandschuricus Schedl, 1941

claviger Blandford, 1894 – KHAB, PRIM.

= platystylus Wichmann, 1915

dahuricus Chapuis, 1869 – CHT, AMUR, PRIM, SAKH, KUR.

= possyeti Stark, 1938

esuriens Blandford, 1894 – KHAB, PRIM, SAKH, KUR.

= sachalinensis Michalski, 1964

jacobsoni (Spessivtsev, 1919) – KHAB, PRIM, SAKH.

= rimskii Kurenzov, 1941

japonicus Chapuis, 1875 – BUR, CHT, KHAB, PRIM.

= confusus Eggers, 1922

= mandli Eggers, 1922

= starki Kurenzov, 1941

= subconfusus Eggers, 1941

= ussuriensis Kurenzov, 1941

kirschii Skalitzky, 1876 – ALT.

= fasciatus Reitter, 1890

koltzei Reitter, 1894 – KHAB, PRIM.

= vexator Reitter, 1913

kononovi Kurenzov, 1941 – PRIM.

mali (Bechstein, 1805) – KURG, BUR, “Western Siberia” [Stark 1952], “Far East” [Knížek 2011].

= pruni Ratzeburg,1837)

= pyri Ratzeburg, 1837

= bicallosus Eggers, 1933

morawitzi Semenov, 1902 – NOV, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

= pini Eggers, 1942

nunbergi Michalski, 1964 – PRIM.

pubescens Stark, 1936 – PRIM.

ratzeburgi Janson, 1856 (Figs. 2g-h) – KHM, TMN, OMS, NOV, TOM, ALT, KEM, KRN, TUV, IRK, BUR, CHT, YAK, KAM, AMUR, KHAB, PRIM, SAKH.

= amurensis Eggers, 1908

= sahlbergi Eggers, 1912

= sibiricus Eggers, 1922

= lineatus Kurenzov, 1941

rugulosus (Mueller, 1818) – KURG, TOM, ALT, “Transbaicalia” [Yanovskij 1999]

= mediterraneus Eggers, 1922

= caucasicus Butovitsch, 1929

= rugulosus samarkandicus Butovitsch, 1929

= manglissiensis Lezhava, 1940

= taxicola Lezhava, 1941

= rugulosus intermedius Sokanovskii, 1960

schevyrewi Semenov, 1902 – NOV, ALT, KHA, KRN, IRK, BUR, CHT, PRIM.

= transcaspicus Eggers, 1922

=sinensis Eggers, 1910

scolytus (Fabricius, 1775) – IRK, “Western Siberia” [Yanovskij 1999].

= destructor Olivier, 1795

= fuchsi Reitter, 1913

semenovi (Spessivtsev, 1919) – BUR, KHAB, PRIM.

ventrosus Schevyrew, 1890 – PRIM, SAKH, KUR, “East Siberia” [Petrov et al. 2019].

= grandis Kurenzov, 1941

= ventricosus Schevyrew, 1897

= trispinosus Strohmeyer, 1908

Tribe Scolytoplatypodini Blandford, 1893

Genus Scolytoplatypus Schaufuss, 1891

daimio Blandford, 1893 (Fig. 3a) – SAKH, KUR.

mikado Blandford, 1893 (Figs. 2i-j) – SAKH, KUR.

tycon Blandford, 1893 (Figs. 2k-l) – AMUR, KHAB, PRIM, SAKH, KUR.

Figure 2.Representatives of Scolytidae: a – Hylastes cunicularius, b – Alniphagus costatus, c – Hylesinus tristis, d – Longulus elatus, e – Dendroctonus micans, f – Pseudohyorrynchus wadai, g – Scolytus ratzeburgi, female, h – S. ratzeburgi, male, i – Scolytoplatypus mikado, male, j – S. mikado, female, k – S. tycon, male, l – S. tycon, female. Photos from www.zin.ru/Animalia/Coleoptera. Author K.V. Makarov.

Tribe Micracidini LeConte, 1876

Genus Pseudothysanoes Blackman, 1920

= Gretschkinia Sokanovskii, 1958

modestus (Murayama, 1940) – “Transbaicalia” [Krivolutskaya 1996].

= mongolica Sokanovskii, 1958

Tribe Ipini Bedel, 1888

Genus Ips De Geer, 1775

acuminatus (Gyllenhal, 1827) – TMN, NOV, TOM, KEM, ALT, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM.

amitinus (Eichhoff, 1871) – TOM, KEM.

duplicatus (Sahlberg, 1836) – KHM, TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

hauseri Reitter, 1894 – KRN, “South Alta” [Stark 1952].

= ussuriensis Reitter, 1913

sexdentatus (Boerner, 1767) – KHM, KURG, TMN, NOV, TOM, KEM, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM.

subelongatus (Motschulsky, 1860) – NOV, KEM, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, KAM, CHUK, MAG, AMUR, KHAB, PRIM, SAKH, KUR.

typographus (Linnaeus, 1758) (Fig. 3b) – KHM, TMN, NOV, TOM, KEM, ALT, RAL, KRN, TUV, IRK, BUR, CHT, YAK, MAG, AMUR, KHAB, PRIM, SAKH, KUR.

Genus Orthotomicus Ferrari, 1867

golovjankoi Pjatnitzky, 1930 – CHT, AMUR, KHAB, PRIM, SAKH.

laricis (Fabricius, 1792) – TMN, TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, MAG, KAM, AMUR, KHAB, PRIM, SAKH, KUR.

proximus (Eichhoff, 1867) – TMN, NOV, TOM, KEM, ALT, RAL, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, PRIM.

starki Spessivtsev, 1926 – RAL, KRN, BUR, KAM, KHAB, PRIM, SAKH.

suturalis (Gyllenhal, 1827) – YAN, KHM, TMN, NOV, TOM, KEM, ALT, RAL, KRN, TUV, IRK, BUR, CHT, YAK, MAG, AMUR, KHAB, PRIM, SAKH, KUR.

= nigritus Gyllenhal, 1827

Genus Pityogenes Bedel, 1888

bidentatus (Herbst, 1784) – NOV, TOM, ALT, RAL, KEM, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH, KUR.

chalcographus (Linnaeus, 1761) – KHM, TMN, OMS, NOV, TOM, ALT, RAL, KRN, TUV, IRK, BUR, CHT, YAK, MAG, KHAB, PRIM, SAKH, KUR.

conjunctus (Reitter, 1887) – TMN, TOM, RAL, KRN, TUV, IRK, BUR, CHT, MAG, PRIM, KUR.

= baicalicus Eggers, 1933

foveolatus Eggers, 1926 – BUR, CHT, MAG, KAM, AMUR, KHAB, PRIM, SAKH, KUR.

irkutensis Eggers, 1910 – TMN, TOM, ALT, RAL, KRN, IRK, BUR, CHT, YAK, KAM, AMUR, PRIM.

= monacensis Fuchs, 1911

quadridens (Hartig, 1834) – TOM, ALT, RAL, KRN, IRK, BUR, CHT, YAK, AMUR.

rudnevi Sokanovskii, 1959 – PRIM.

saalasi Eggers, 1914 – RAL, KRN, IRK, BUR, YAK.

seirindensis Murayama, 1929 – KHAB, PRIM, SAKH.

= aizawai Kono, 1938

= nitidus Eggers, 1941

Genus Pityokteines Fuchs, 1911

curvidens (Germar, 1824) – YAK.

Tribe Dryocoetini Lindemann, 1877

Genus Dryocoetes Eichhoff, 1864

aceris Krivolutskaya, 1968 – KUR.

alni (Georg, 1856) – RAL, KRN, IRK.

autographus (Ratzeburg, 1837) (Fig. 3c) – YAN, TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, MAG, AMUR, KHAB, PRIM, SAKH, KUR.

= suecicus Eggers, 1923

baikalicus Reitter, 1899 – NOV, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, MAG, AMUR, KHAB, SAKH, KUR.

infuscatus Murayama, 1937 – BUR, CHT, PRIM, SAKH, KUR.

= orientalis Kurenzov, 1941

= orientalis var. pilosiusculus Kurenzov, 1948

hectographus Reitter, 1913 – KHM, TOM, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH, KUR.

krivolutzkajae Mandelshtam, 2001 – KAM.

pini Niisima, 1909 – BUR, CHT, SAKH.

cerasi Eggers, 1942 – PRIM.

=cerasi Stark, 1950

padi Kurenzov, 1941 – KHAB, PRIM.

= padi Stark, 1952

rugicollis Eggers, 1926 – AMUR, KHAB, PRIM, SAKH, KUR.

striatus Eggers, 1933 – PRIM, SAKH, KUR.

= abietinus Kono et Tamanuki, 1939

uniseriatus Eggers, 1926 – SAKH.

ussuriensis Eggers, 1933 – MAG, KAM, AMUR, KHAB, PRIM, SAKH, KUR, “Western Siberia ” [Knížek 2011].

= rugulosus Eggers, 1933

Genus Lymantor Lovendal, 1889

aceris (Lindemann, 1875) – BUR, PRIM.

coryli (Perris, 1853) – IRK, BUR, YAK, “Western Siberia, Far East” [Alonso-Zarazaga et a. 2017].

Genus Taphrorychus Eichhoff, 1878

carpini (Kurenzov, 1941) – PRIM.

= carpini Eggers, 1942

= carpini Stark, 1952

Tribe Crypturgini LeConte, 1876

Genus Crypturgus Erichson, 1836

cinereus (Herbst, 1793) – NOV, TOM, KEM, ALT, RAL, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

hispidulus Thomson, 1870 – TOM, KEM, RAL, KRN, BUR, CHT, YAK, KAM, PRIM, SAKH, KUR.

pusillus (Gyllenhal, 1813) – TOM, KEM, RAL, KRN, IRK, BUR, CHT, YAK, PRIM, SAKH, KUR.

subcribrosus Eggers, 1933 – KRN, PRIM, SAKH, “Eastern Siberia” [Knížek 2011].

tuberosus Niisima, 1909 – KHAB, PRIM, SAKH.

Tribe Xyloterini LeConte, 1876

Genus Indocryphalus Eggers, 1939

aceris (Niisima, 1910) (Fig. 3d) – KHAB, PRIM.

pubipennis (Blandford, 1894) – “SAKH, KUR” [Krivolutskaya 1996].

Genus Trypodendron Stephens, 1830

laeve Eggers, 1939 – KRN.

lineatum (Olivier, 1795) – KHM, TMN, TOM, NOV, KEM, ALT, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, MAG, KAM, AMUR, KHAB, PRIM, SAKH, KUR.

=granulatum Eggers, 1933

niponicum Blandford, 1894 – BUR, MAG, KAM, KHAB, PRIM, SAKH, KUR.

proximum Niisima, 1909 (Fig. 3e) – KHAB, PRIM, SAKH, KUR.

signatum (Fabricius, 1787) – NOV, KEM, RAL, KRN, TUV, IRK, BUR, CHT, YAK, PRIM, SAKH.

=suturale Eggers, 1933

Tribe Xyleborini LeConte, 1876

Genus Anisandrus Ferrari, 1867

apicalis Blandford, 1894 – PRIM, KUR (Kunashir).

dispar (Fabricius, 1792) (Figs. 3g-h) – TMN, TOM, OMS, NOV, KEM, ALT, RAL, KHA, KRN, TUV, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH.

= aequalis Reitter, 1913

maiche Stark, 1936 –PRIM, KUR, “Western Siberia” [Krivolutskaya 1996], “Eastern Siberia” [Knížek 2011].

Genus Cnestus Sampson, 1911

mutilatus (Blandford, 1894) – PRIM.

Genus Cyclorhipidion Hagedorn, 1912

bodoanus Reitter, 1913 – EAO, PRIM. “Eastern Siberia” [Stark 1952].

= punctulatus Kurenzov, 1948

japonicus (Nobuchi, 1981) – PRIM.

pelliculosum (Eichhoff, 1878) – PRIM.

=starki Nunberg, 1956

= quercus Kurenzov, 1948

Genus Microperus Wood,1980

molestus Park et Smith, 2020 – PRIM.

Genus Xyleborinus Reitter, 1913

attenuatus (Eichhoff, 1876) – KHAB, PRIM, SAKH.

= alni Niisima, 1909

saxesenii (Ratzeburg, 1837) – TUV, BUR, YAK, KAM, PRIM, SAKH, KUR, “Western Siberia” [Knížek 2011].

Figure 3.Representatives of Scolytidae: a – Scolytoplatypus daimio, male, b – Ips typographus, c – Dryocoetes autographus, d – Indocryphalus aceris, e – Trypodendron proximum, f – Xyleborus seriatus, g – Anisandrus dispar, female, h – A. dispar, male, i – Ernoporicus insularum. Photos from www.zin.ru/Animalia/Coleoptera. Author K.V. Makarov.

Genus Xyleborus Eichhoff, 1864

cryptographus (Ratzeburg, 1837) – KRN, “Altai” [Yanovskij 1999], “Western Siberia, Far East” [Knížek 2011].

seriatus Blandford, 1894 (Fig. 3f) – PRIM.

= orientalis Eggers, 1933

Genus Xylosandrus Reitter, 1913

germanus (Blandford, 1894) – PRIM, SAKH, KUR.

Tribe Cryphalini Lindemann, 1877

Genus Allernoporus Kurenzov, 1941

euonymi Kurenzov, 1941 – PRIM.

Genus Cryphalus Erichson, 1836

asperatus (Gyllenhal, 1813) – TOM, KRN, IRK, “Far East” [Knížek 2011].

= abietis Ratzeburg, 1837

carpini Berger, 1916 – PRIM.

= carpinivorus Murayama, 1930

coryli Stark, 1936 – PRIM.

exiguus Blandford, 1894 – SAKH, KUR.

kurenzovi Stark, 1936 – AMUR, PRIM, SAKH, KUR.

= punctulatus Eggers, 1942

= ussuriensis Eggers, 1942

kurilensis Krivolutskaya, 1968 – KUR.

laricis Niisima, 1909 – SAKH.

latus Eggers, 1929 – BUR, CHT, YAK, AMUR, KHAB, PRIM, SAKH, “Western Siberia” [Alonso-Zarazaga et a. 2017].

= premayaensis Murayama, 1943

longus (Eggers, 1926) – KAM, MAG, PRIM, SAKH, KUR.

= alni Krivolutskaya, 1958

malus Niisima, 1909 – SAKH, KUR.

= padi Krivolutskaya, 1958

mandschuricus Eggers, 1929 – KHAB, PRIM.

nataliyae Mandelshtam et Petrov, 2022 – SAKH.

piceae (Ratzeburg, 1837) – KHAB, PRIM.

= orientalis Eggers, 1911

piceus Eggers, 1926 – KHAB, PRIM, SAKH, KUR.

pruni Eggers, 1929 – PRIM, SAKH.

redikorzevi Berger, 1916 – PRIM, SAKH, KUR.

saltuarius Weise, 1891 – RAL, KRN, IRK, BUR, CHT, YAK, AMUR, KHAB, PRIM.

scopiger Berger, 1916 – PRIM.

sichotensis Kurenzov, 1941 – KHAB, PRIM.

viburni Stark, 1936 – PRIM.

= viburni Eggers, 1942

Genus Eidophelus Eichhoff, 1875

imitans Eichhoff, 1875 – PRIM, SAKH, KUR.

= elegans Krivolutskaya, 1958

Genus Ernoporicus Berger, 1917

corni (Kurenzov, 1941) – KHAB, PRIM, SAKH.

insularum (Krivolutskaya, 1968) (Fig. 3i) – KUR.

= krivolutskayae Wood, 1992

semenovi (Kurenzov, 1941) – PRIM, SAKH, KUR.

spessivtzevi Berger, 1916 – PRIM, SAKH, KUR.

zachvatkini (Krivolutskaya, 1958) – SAKH, KUR.

Genus Ernoporus Thomson, 1859

tiliae (Panzer, 1793) – PRIM, “Western Siberia” [Stark 1952].

= eggersi Stark, 1936

= starki Eggers, 1942

Genus Hypothenemus Westwood, 1834

atomus Hopkins, 1915 – PRIM.

margaritae Petrov and Shamaev, 2020 – PRIM.

Genus Procryphalus Hopkins, 1915

fraxini (Berger, 1916) – KHAB, PRIM.

Genus Trypophloeus Fairmaire, 1868

alni (Lindemann, 1875) – RAL, KRN, IRK, MAG, KAM, PRIM, SAKH.

binodulus (Ratzeburg, 1837) – NOV, KRN, ALT, RAL, PRIM, SAKH.

= berezinae Stark, 1952

=kurenzovi Nunberg, 1956

= kurenzovi Schedl, 1959

= populi Kurenzov, 1941

=grothii Hagedorn, 1904

bispinulus Eggers, 1927 – RAL, BUR, PRIM.

dejevi Stark, 1936 – BUR, KAM, MAG, PRIM, SAKH, “Western Siberia” [Knížek 2011].

= dejevi Eggers, 1942

= niger Stark, 1936

Tribe Corthylini LeConte, 1876

Subtribe Pityophthorina Eichhoff, 1878

Genus Pityophthorus Eichhoff, 1864

abietinus Wood, 1989 – PRIM.

= abietis Kurenzov, 1941

= kurentzovi Krivolutskaya, 1996

glabratus Eichhoff, 1878 – RAL, KRN.

jucundus Blandford, 1894 – SAKH.

lapponicus Kurenzov, 1941 – PRIM, “Eastern Siberia” [Yanovskij 1999].

= lapponicus Stark, 1952

lichtensteinii (Ratzeburg, 1837) – KEM, RAL, KRN, IRK, YAK, AMUR, KUR.

= rossicus Eggers, 1915

micrographus (Linnaeus, 1758) – RAL, KRN, IRK, BUR, YAK, AMUR.

morosovi Spessivtsev, 1926 – RAL, KRN, IRK, BUR.

pini Kurenzov, 1941 – KRN, IRK, YAK, PRIM.

sachalinensis Krivolutskya, 1956 – SAKH.

sichotensis Kurenzov, 1941 – KEM, IRK, PRIM.

traegardhi Spessivtsev, 1921 – RAL, KRN, IRK, YAK, “Far East” [Knížek 2011].

Subtribe Corthylina LeConte, 1876

Genus Monarthrum Kirsch, 1866

meuseli (Reitter, 1905) – KRN.

Family Platypodidae Shuckard, 1840

Subfamily Platypodinae Shuckard, 1840

Tribe Platypodini Shuckard, 1840

Genus Platypus Herbst, 1793

koryoensis (Murayama, 1930) – PRIM.

quercivorus (Murayama, 1925) – PRIM.

Genus Treptoplatypus Schedl, 1972

severini (Blandford, 1894) (Fig. 4) – PRIM, KUR.

Currently, 185 species of the family Scolytidae and three species of the family Platypodidae are recorded from Asian Russia, 99 species of bark beetles are found in Siberia and 168 species in the Russian Far East. Platypodidae are known from the of the the Russian Far East. Two species are found in Yamalo-Nenets Autonomous Okrug, 13 species in Khanty-Mansi Autonomous Okrug, 19 species in Tyumen Oblas, four species in Kurgan Oblast, three species in Omsk Oblast, 37 species in Tomsk Oblast, 28 species in Novosibirsk Oblast, 32 species in Kemerovo Oblast, 25 species in Altay Krai, 53 species in Altai Republic, 62 species in Krasnoyarsk Krai, 11 species in Republic of Khakassia, 24 species in Tyva Republic, 55 species in Irkutsk Oblast, 61 species in Buryatiya Republic, 40 species in Zabaikalskii Krai, 52 species in Sakha (Yakutia) Republic, 25 species in Kamchatka Oblast, one species in Chukotka Autonomous Okrug, 19 species in Magadan Oblast, 45 species in Amur Oblast, one species in Jewish Autonomous Oblast, 67 species in Khabarovsk Krai, 130 species in Primorsky Krai, 87 species in Sakhalin Is. and 68 species in Kuriles Isl.

Figure 4.Treptoplatypus severini: a – male, b – female. Photos from www.zin.ru/Animalia/Coleoptera. Author K.V. Makarov.

Acknowledgements

The author thanks V.V. Dubatolov (Novosibirsk), R.Yu. Dudko (Novosibirsk), D.A. Efimov (Kemerovo), A.V. Korshunov (Kemerovo), S.V. Reshetnikov (Novosibirsk), E.Yu. Shevnin (Novosibirsk), S.E. Tshernyshev (Novosibirsk), V.K. Zinchenko (Novosibirsk) for the opportunity to study material.

References

Agafonova TA, Silaev AS, Antonov IA (2021) Bark beetles (Coleoptera, Curculionidae: Scolytinae) of the Baikal Siberia. Izvestiya Irkutskogo gosudarstvennogo universiteta. Seriya Biolo-giya. Ekologiya 37, 54–69. https://doi.org/10.26516/2073-3372.2021.37.54 [In Russian].

Akulov EN, Mandelshtam MYu (2012a) New data on scolytid fauna (Coleoptera, Curculionidae, Scolytinae) of the Central Siberia. XIV Congress of the Russian Entomological Society. Sankt-Peterburg. P. 13 and poster. https://doi.org/10.13140/RG.2.2.23029.12007 [In Russian].

Akulov EN, Mandelstam MYu (2012b) About new finds of bark beetles (Coleoptera: Сurculionidae: Scolytinae) in the south of the Krasnoyarsk Territory and in the Republic of Khakassia. Ecological and economic consequences of invasions of dendrophilic insects. Materials of the All-Russian conference with international participation. Krasnoyarsk: IL SO RAN, 123–128. [In Russian].

Alonso-Zarazaga MA, Barrios H, Borovec R, Bouchard P, Caldara R, Colonnelli E, Gültekin L, Hlavá P, Korotyaev B, Lyal CHC, Machado A, Meregalli M, Pierotti H, Ren L, Sánchez-Ruiz M, Sforzi A, Silfverberg H, Skuhrovec J, Trýzna M, Velázquez de Castro AJ, Yunakov NN (2017). Cooperative catalogue of Palaearctic Coleoptera Curculionoidea. Monografías electrónicas 8, 1–729.

Atlas of beetles of Russia: Superfamily Curculionoidea. https://www.zin.ru/Animalia/Coleoptera/rus/atl_cur.htm

Averensky AI (1971) On the fauna of bark beetles (Coleopters, Ipidae) of coniferous species of southwestern Yakutia. Harmful insects and helminths of Yakutia. Yakutsk: YaFSOAN SSSR, 12–16. [In Russian].

Averensky AI (1999) The catalogue of beetles of Yakutia. Yakutsk, Yakustk Scientific Centre SD RAS. 77 pp. [in Russian].

Berezhnykh ED (1979) Xylophagous insects of the Muya basin. Animal parasites and plant pests of the Baikal and Transbaikalia. Ulan-Ude: BFSOANSSSR, 12–16. [In Russian].

Bukhkalo SP, Galich DE, Sergeeva EV, Alemasova NV (2011). Synopsis of beetle fauna of the southern taiga of Western Siberia (lower of Irtysh basin). KMK, Moscow, 267 pp. [In Russian].

Bukhkalo SP, Galich DE, Sergeeva EV, Vazhenina NV (2014) Synopsis of invertebrate fauna of the southern taiga of Western Siberia (lower of Irtysh basin). KMK, Moscow, 189 pp. [In Russian].

Cherepanov AI (1952) Harmful insects of field-protective forest belts. Novosibirsk: OGIZ, 128 pp. [In Russian].

Efimov DA, Legalov AA (2012) New records of the Curculionoid beetles (Coleoptera) from Kuznetsk-Salair mountain area. Amurskii Zoologicheskii Zhurnal 4 (3), 247–249. [In Russian].

Eggers H (1942) Zur palaearktischen Borkenkäferfauna. (Coleoptera: Ipidae). VIII. Borkenkäfer aus dem asiatischen Russland. Arbeiten über Morphologische und Taxonomische Entomologie 9, 27–36.

Frolov DN (1949) Bark beetles of coniferous trees of Eastern Siberia. Irkutsk: OGIZ, 138 pp. [In Russian].

Galkin GI (1966) Insects - pests of pine plantations in the Angara region. Trudy SibNIILP, 14, 82–93.

Gratshev VG, Legalov AA (2014) The Mesozoic stage of evolution of the family Nemonychidae (Coleoptera, Curculionoidea). Paleontological Journal, 48, 851–944. https://doi.org/10.1134/S0031030114080012.

Gurov AV, Dryannykh NM (1982) Phytophageous insects in pine forests of the northern part of the Krasnoyarsk forest-steppe. Insects of forest-steppe forests. Novosibirsk: Nauka, 5–18. [In Russian].

Heyden L (1880–1881) Catalog den Coleopteren von Sibirien mit Einschluss derjenigen der Turanischen Länder, Turkestans und der chinensischen Grenzgebiete. Berlin, 224 pp.

Isaev AS, Tarasova DA (1965) Stem pests of pine in the Middle Amur Region and Research on the protection of Siberian forests. M.: Nauka, 5–19. [In Russian].

Ivliev LA, Kononov DG (1966) New data on bark beetles (Coleoptera, Ipidae) of the Magadan region. Harmful insects of the forests of the Soviet Far East. Vladivostok, 5–42. [In Russian].

Ivliev LA, Kononov DG (1974) New data on bark beetles (Coleoptera, Ipidae) of Sakhalin. Proceedings of the seventh congress of the VEO, 2. Leningrad, 214–215. [In Russian].

Izhevsky SS, Nikitsky NB, Volkov OG, Dolgin MM (2005) An illustrated reference book of xylophagous beeles damaging forests and timber in Russia. Tula: Grif, 218 pp. [In Russian].

Johnson AJ, Li Y, Mandelshtam MYu, Park S, Lin C-S, Gao L, Hulcr J (2020) East Asian Cryphalus Erichson (Curculionidae, Scolytinae): new species, new synonymy and redescriptions of species. ZooKeys 995: 15–66. https://doi.org/10.3897/zookeys.995.55981

Kerchev IA (2011) Species diversity of xylophagous insects in the southern taiga Siberian pine forests. Conceptual and applied aspects of research and education in invertebrate zoology. Romanenko VN (ed.). Tomsk: Agraf-Press, 62–66. [In Russian].

Kerchev IA, Mandelshtam MYu, Krivets SA, Ilinsky YuYu (2019) Small spruce bark beetle Ips amitinus (Eichhoff, 1872) (Coleoptera, Curculionidae: Scolytinae): a new alien species in West Siberia. Entomological Review 99, 639–644.

Kiseleva EF (1928) A note about bark beetles in the environs of Tomsk. Izvestiya Tomskogo Universiteta 79 (4), 243–246. [In Russian].

Kiseleva EF (1946) Bark beetles of Tomsk Province. Trudy Tomskogo Gosudarstvennogo Universiteta 97, 123–136. [In Russian].

Kiseleva EF (1951) Pests of the siberian pine and methods of their control in Tomsk Province. Uchenye Zapiski Tomskogo Universiteta 15, 85–100. [In Russian].

Kiseleva EF (1952) A review of insect pests of Tomsk Province and methods of their control. Trudy Tomskogo Gosudarstvennogo Universiteta 118, 47–60. [In Russian].

Knížek M (2011) Scolytinae and Platypodinae. Catalogue of Palaearctic Coleoptera.7. Löbl I., Smetana A. (eds.). Stenstrup: Apollo Books, 86–87, 204–251.

Kononenko AP, Opanasenko FI (1971) The role of stem pests of coniferous trees in the mountain forests of the North-Eastern Altai. Protection of the forest from harmful insects and diseases, 3. M., 57–9. [In Russian].

Krivets SA, Chemodanov AV (2005) Insect forest pests in Tomsk Province. Entomological Studies in Siberia, 4 (Baranchikov YuN, ed.). Krasnoyarsk, 98–118. [In Russian].

Krivets SA, Kerchev IA, Bisirova EM, Demidko DA, Pet'ko VM, Baranchikov YuN (2011) Distribution of four-eyed fir bark beetle Polygraphus proximus Blandf. (Coleoptera, Curculionidae: Scolytinae) in Siberia. Izvestiya Sankt-Peterburgskoy lesotekhnicheskoy akademii 211, 33–45. [In Russian].

Krivets SA, Vysotina SV (2011) Ecological and faunistic review of the Coleopterous (Insecta, Coleoptera) dendrophagans of the priketye conifer forests (Tomsk Oblast). Vestnik Tomskogo gosudarstvennogo universiteta. Biologiya 3 (15), 111–126. [In Russian].

Krivolutskaya GO (1958) Bark beetles of Sakhalin Island. Moskva-Leningrad: Akademia nauk SSSR, 196 pp. [In Russian].

Krivolutskaya GO (1961) Pests of afforestation in Novosibirsk. Plant resources of Novosibirsk Province. Novosibirsk, 199–207. [In Russian].

Krivolutskaya GO (1965a) Cryptic trunk pests in West Siberian dark coniferous forests damaged by the Siberian moth. Moscow: Nauka, 130 pp. [In Russian].

Krivolutskaya GO (1965b) Fauna of bark beetles (Coleoptera, Ipidae) of the southern Kuril Islands. Forestry research in the Far East. Vladivostok: DVFSOANSSSR, 219–243. [In Russian].

Krivolutskaya GO (1973) Entomofauna of Kuril Islands. L.: Nauka, 315 pp. [In Russian].

Krivolutskaya GO (1983) Ecological-geographical characteristics of the fauna of bark beetles of Northern Asia]. Entomologicheskoe Obozrenie 62 (2), 287–301. [In Russian].

Krivolutskaya GO (1996a) Family Scolytidae — bark-beetles. Keys to the insects of the Russian Far East. 3(3). Lehr P.A. (ed.). Vladivostok: Dal’nauka, 312–373. [In Russian].

Krivolutskaya GO (1996b) Family Platypodidae. Keys to the insects of the Russian Far East. 3(3). Lehr P.A. (ed.). Vladivostok: Dal’nauka, 374–377. [In Russian].

Krivolutskaya GO, Kupyanskaya AN (1970) Bark beetles (Coleoptera, Ipidae) in urban green spaces of Primorsky Krai. Entomological research in the Far East. Vladivostok, 185–195. [In Russian].

Kurentsov AI (1941) Bark-beetles of the USSR Far East. Moscow-Leningrad: Izdatel’stvo Akademii Nauk SSSR, 234 pp. [In Russian].

Lavrov CD (1927) Materials to studying of entomophauna env. of Omsk. Proceedings of Siberian institute of agriculture and forestry. Omsk 8 (3), 51– 100. [In Russian].

Legalov AA (2015) Fossil Mesozoic and Cenozoic weevils (Coleoptera, Obrienioidea, Curculionoidea). Paleontological Journal , 49, 1442–1513, https://doi.org/10.1134/S0031030115130067.

Legalov AA, Sitnikov PS (2000) Materials on the fauna weevils-beetles (Coleoptera, Curculionoidea) of Tyumen Area. Vestnik ekologii, lesovedenija i landshaftovedenija. Tyumen 1, 37–47 [In Russian].

Lindeman GV (1961) On the fauna and ecology of bark beetles in Southwestern Transbaikalia. Communications of the forestry laboratory, 4, 98–101. [In Russian].

Lurie MA, Lindeman GV (1961) Stem pests of Dahurian larch in Transbaikalia. Izvestiya SO AN USSR, 2, 116–120. [In Russian].

Mandelshtam MJu (2001) A new species of bark-beetles (Coleoptera: Scolytidae) from Russian Far East. Far Eastern Entomologist 105, 11–12.

Mandelshtam MJu (2002). New synonymy, new records and lectotype designation in Palaearctic Scolytidae (Coleoptera). Far Eastern entomologist 119, 6–11.

Mandelshtam MYu, Petrov AV (2016) Notes on Oriental Scolytus (Coleoptera: Curculionidae: Scolytinae). Zoosystematica Rossica 25 (2), 295–298.

Mandelshtam MYu, Petrov AV (2019) A key to species of the tribe Hylastini LeConte, 1876 (Coleoptera: Curculionidae: Scolytinae) from Russia and adjacent countries. Russian Entomological Journal 28 (4), 389–399. https://doi.org/10.15298/rusentj.28.4.08

Mandelshtam MYu, Petrov AV (2022) A new Cryphalus Erichson, 1836 species (Coleoptera: Curculionidae: Scolytinae) from Sakhalin. Russian Entomological Journal 31 (2), 159–162. https://doi.org/10.15298/rusentj.31.2.11

Mandelshtam MYu, Petrov AV, Barclay MVL, Knížek M, Beaver RA (2007) Taxonomic changes in Scolytinae (Coleoptera: Curculionidae: Scolytinae) from Eastern Asia. Russian Entomological Journal 16 (1), 459–464.

Mandelshtam MYu, Sergeev ME (2020) New records of bark beetles (Coleoptera, Curculionidae: Scolytinae) in Sikhote-Alin Nature Reserve. Dendrobiontnyye bespozvonochnyye zhivotnyye i griby i ikh rol' v lesnykh ekosistemakh. Sankt-Peresburg, 216–217. [In Russian].

Mandelshtam MYu, Yakushkin EA, Kovalenko YaN, Petrov AV (2022) New invasive Hypothenemus Westwood, 1834 (Coleoptera, Curculionidae: Scolytinae) species in the Caucasus and in the Southern Primorskii Territory, Russia. Entomological Review 102 (4), 485–491. https://doi.org/ 10.1134/S0013873822040078

Mandelshtam MYu, Yakushkin EA, Petrov AV (2018) Oriental ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) – new inhabitants of Primorsk territory in Russia. Russian Journal of Biological Invasions 3, 74–87. [In Russian].

Morimoto K, Kojima, H (2004) Systematic position of the tribe Phylloplatypodini, with remarks on the definitions of the families Scolytidae, Platypodidae, Dryophthoridae and Curculionidae (Coleoptera: Curculionoidea). Esakia, 44, 153–168.

Nemkova RI (1965) Stem pests of dark coniferous forests of the Western Sayan. Krasnoyarsk: Krasnoyarsk kizhnoe izdatel’stvo. 87 pp. [In Russian].

Nobuchi A (1979) Studies on Scolytidae XVIII. Bark beetles of tribe Polygraphini (Coleoptera, Scolytidae) in Japan. Bulletin of Forestry and Forest Products Research Institute 308, 1–16.

Opanasenko FI, Kononenko AP (1966) Species composition and ecological features of stem insects - inhabitants of the fir of the North-Eastern Altai. Fauna and ecology of arthropods of Siberia. Novosibirsk: Nauka, 83–86. [In Russian].

Park S, Lyu D (2007) Checklist of the Family Platypodidae (Coleoptera) in Korea. Journal of Asia-Pacific Entomology 10 (3), 275–280, https://doi.org/10.1016/S1226-8615(08)60363-5

Petrov AV (2011) New data on bark beetles of the genus Hylesinus Fabricius, 1801 from Russia and adjacent countries. Lesnoy Vestnik. 4(80), 21–24 [In Russian].

Petrov AV (2018) A key to genera and species of the tribe Hylesinini Erichson, 1836 (Coleoptera: Curculionidae: Scolytinae) from Russia and adjacent countries. Russian Entomological Journal 27 (2), 179–189. https://doi.org/10.15298/rusentj.27.2.08

Petrov AV, Mandelshtam MJu (2002) New synonymy in the genus Hylesinus Fabricius (Coleoptera: Scolytidae). Far Eastern Entomologist 119, 11–12.

Petrov AV, Mandelshtam MYu, Beaver RA (2019) A key to species of the tribe Scolytini Latreille, 1804 (Coleoptera: Curculionidae: Scolytinae) from Russia and adjacent countries. Russian Entomological Journal 28 (3), 286–302. https://doi.org/10.15298/rusentj.28.3.08

Petrov AV, Shamaev AV (2020) Description of a new Hypothenemus Westwood, 1834 species (Coleoptera: Curculionidae: Scolytinae) from South of Primorskiy Kray of Russia and South Korea. Russian Entomological Journal 29 (1), 83–86. https://doi.org/10.15298/rusentj.29.1.11

Rudnev DF (1958) Bark beetles (Coleoptera, Ipidae) of the Magadan region. Entomologicheskoe Obozrenie, 37 (2), 369–373. [In Russian].

Sokanovsky BV (1960) On the taxonomy and distribution of bark beetles (Coleoptera, Ipidae) in the USSR and adjacent countries. Entomologicheskoe Obozrenie, 39 (3), 674–678. [In Russian].

Shamaev AV (2021) Illustrated key to bark beetle species of the genus Ips DeGeer (Coleoptera: Curculionidae, Scolytinae) of Russia based on their damage to the bark. Euroasian Entomological Journal, 20 (6), 299–306. https://doi.org/10.15298/euroasentj.20.6.01

Shatilov OA (1985) On the biology of little-known species of bark beetles (Coleoptera, Ipidae) of Altai. Systematics and biology of arthropods and helminths. Novosibirsk: Nauka, 54–56. [In Russian].

Shatilov OA (1987) On the study of the vertical distribution of bark beetles (Ipidae) in Altai. Ecology and geography of arthropods of Siberia. Novosibirsk: Nauka, 120–122. [In Russian].

Stark VN (1952) Bark beetles. Fauna SSSR. Zhestkokrylye. 31. Moscow–Leningrad: Izdatel’stvo Akademii Nauk SSSR. 462 pp. [In Russian].

Wood SL (1986) A reclasification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs, 10, 1–126.

Wood SL (1993) Revision of the genera of Platypodidae (Coleoptera). Great Basin Naturalist Memoirs, 53, 259–281.

Wood SL, Bright DE (1992) A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic Index. Great Basin Naturalist Memoirs, 13, 1–1553.

Yanovskij VМ (1999) Annotated list of scolytids (Coleoptera, Scolytidae) of North Asia. Entomologicheskoe obozrenie 78 (2), 327–362. [In Russian].

Zherichin VV, Egorov AB (1991) Weevils (Coleoptera, Curculionidae) from Russian Far East (review of subfamilies with description new taxa). Vladivostok, 164 pp. [In Russian].